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<br />Southwestern Willow Flycatcher <br /> <br />By: Debra T. Bills, U.S. Fish and Wildlife Service <br />Bryan T. Brown, Consulting Ecologist <br /> <br />II <br /> <br />The Southwestern willow flycatcher (Empidonax traillii extimus) is a rare riparian bird fOWld in Arizona, Ne\'" <br />Mexico, and southern California. The extimlJ.s subspecies of willO\\' flycatcher is rarest in Arizona. In 1987, <br />approximately 2S pairs of willow flycatchers were believed to he in Arizona, \\;th the largest population-7 pairsnin <br />the Grand Canyon. <br /> <br />A 1991 slln'ey of ....illow flycatchers below Glen Canyon Dam in the Glen Canyon National Recreation Area and <br />Grand Canyon National Park located only two nesting pairs. The most probable reason for the apparent decline <br />in the Grand Canyon is brood parasitism by brown-headed cowbirds. Instead of making their own nests, <br />brown-headed cowbirds lay their eggs in other species nests, usually at the expense of their host's young. If the <br />host's young or eggs are not pushed out of the nest by the adult cowbirds, the young willow flycatchers usually <br />succumb to competitive pressure from the larger, more aggressive young cowbirds. <br /> <br />Compounding the threat of cowbird parasitism is habitat t Y t!~ ~- <br />fragmentation caused by floods and fluctuating flows. Fluctuating r~ ~l ~,.. <br />flows contribute to the erosion of terrestrial habitats, resulting in .. J ~ <br />decreased size of contiguous vegetation patches. Many species ,. -,Jft.. <br />require specific threshold patch sizes for nesting. Although the r. <br />required specific patch size for nesting willow flycatchers is not <br />known, it is known that the brown-headed cowbird prefers small <br />patch sizes and habitats with a high edge to center ratio. <br /> <br />The willow flycatcher is a neotropical migrant songbird which <br />arrives in Arizona in early May. and in the Grand Canyon area <br />around May 15. Nesting begins in late May and continues through <br />July_ The average clutch is three or four eggs, and incubation is <br />complete in about 12 days. The nestlings spend another 12 days or <br />so on the nest. <br /> <br />I" <br />," <br /> <br />Although the willow flycatcher has traditionally been associated <br />with willows and other native riparian vegetation, all of the nests <br />locdted in the Grand Canyon have been located in tamarisk (salt <br />cedar), even though nath'e vegetation was available. <br /> <br />These secretive birds are almost impossible to visually distinguish <br />from other subspecies of willow flycatchers. However, since the <br />southwestern subspecies is the only willow flycatcher known to breed in Arizona, finding a nest or hearing the <br />classic male "fitz bew" song during the breeding season is a good indication that it is the extimus subspecies. <br /> <br />The Arizona Game and Fish Department considers the willow flycatcher as endangered in Arizona. The US Fish <br />and Wildlife Service classifies the willow flycatcher as a Category 1 species. (A species or subspecies for which <br />there is substantial information to support the biological appropriateness of proposing to list it as threatened or <br />endangered.) <br /> <br />If you have any questions or information on this species, please contact Debra Bills or Sam Spiller, Field Supen-isor. <br />U.S. Fish and Wildlife Service, Phoenix, Arizona. (602) 379-4720. <br /> <br /> <br />, <br />Skrklt by Brio" Ero"J <br /> <br />12 <br />