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<br />002036 <br /> <br />for greenbacks, appears to be allcwing for the downstream expansion of <br />cutthroats into brook trout populations in some streams within RMNP (North <br />Fork of the Big Thompson River, North Inlet and North st. Vrain). However, in <br />other areas, (OUzel, Hidden Valley, George and cornelius Creeks) brook trout <br />continue to expand into populations cf greenbacks despite no legal angler <br />harvest of greenbacks. <br /> <br />Habitat Reauirements <br /> <br />Habitat requirements of greenback cutthroat trout appear little different from <br />other species of trout. Bulkley (1959) gathered information on age, growth, <br />food habits, and movement of a slightly hybridized population in the . <br />headwaters (3,200 m) of the Big Thompson River, Rocky Mountain National Park <br />(RMNP). Nelson (1972) provided data on age, growth, and fecundity of a dense, <br />unexploited, and slightly hybridized greenback population in Island Lake, <br />Boulder Creek watershed. <br /> <br />Restoration efforts shculd be directed to habitats that are capable of <br />supporting a minimum of 20 kg/ha of fish. Habitats occupied by non-native <br />fish will require their removal prior to the introduction of greenbacks to <br />prevent hybridization and competition. <br /> <br />Stable reproducing populations of greenbacks in Colorado are rarely found <br />above timberline since cold water temperatures do not allow for sufficient <br />time for spring spawning, hatching and establishment of fry during the short <br />ice-free pericd. Currently, the highest known elevation of a long-term <br />reproducing population is the Upper Hutcheson Lake population at 3,402 m. Due <br />to the availability of surplus greenbacks, experimental introductions are <br />being conducted in high elevation fishless waters tc document the effect of <br />elevation as a limiting factor on greenbacks. Two timberline lakes that were <br />stocked with non-native cutthroats in RMNP, but became fishless after the <br />termination of the non-native fish stocking, were subsequently stocked with <br />native greenbacks. In one of these lakes (Lake Odessa at 3,048 m), greenbacks <br />spawned. from late June to early July and established a reproducing population. <br />Greenbacks stocked in the other lake, Crystal Lake (3,511 m) spawned by mid- <br />July, but survival past the egg stage was not documented through 1992. <br /> <br />The lower elevation limit of greenback survival is not known. However, <br />greenbacks stocked in a low elevation lake (1,889 m) on Fort Carson, Colorado, <br />have survived and attained a size of 2.0 kg. Future experimental stocking <br />should involve lower elevation projects to determine greenback survival in low <br />elevation habitats, and in association with native non-salmonid forage species <br />in low elevation habitats. <br /> <br />ReDroduc~iaD <br /> <br />spawning is generally initiated in the spring when water temperatures reach <br />5C-8C. Due to the influence of elevation on water temperatures, greenbacks in <br />Lytle Pond on Ft. Carsonc(.l,889 m) spawn by early April, greenbacks in Hunters <br />Creek (2,896 m) spawn in mid-June, and greenbacks in Upper Hutcheson Lake <br />(3,402 m) spawn in mid-July. Although greenbacks are spring spawners, older <br />greenback males in high elevation streams (Hunters Creek and the headwaters of <br />the North Fork of the Big Thompson River within RMNP), were observed to be in <br />spawning colors and running milt in mid-September. <br /> <br />Although Como Creek greenbacks can produce eggs at age 2 in the hatchery, <br />females in small subalpine streams within Colorado appear to mature after <br />their third to fourth summer of life when they reach lengths of approximately <br />180 mm. <br /> <br />9 <br />