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at 11 sandpits in 1984 when riverine habitat was unavailable because of high <br />water levels. Additional measurements of colony spatial characteristics during <br />years of contrasting flow conditions would facilitate further evaluation of <br />I this matter. <br />Certain parameters such as area per colony, nest elevation above water level, <br />and nest -to -water distances are subject to change during the breeding season <br />with fluctuations in river stage. All of these parameters vary inversely with <br />river stage and, in general, decreasing stage conditions are more favorable to <br />tern and plover colonies. Increases in'area per colony resulting from stage <br />decreases tend to reduce the potential for predation, whereas associated <br />increases in nest elevation and nest -to -water distances reduce potential <br />disturbances from wave action or subsequent rises in water levels (Schwalbach <br />et al. 1986). Stage increases cause reverse effects and, in the event of large <br />increases, complete inundation of nesting areas can occur. <br />Although several researchers (Ducey 1981b, Faanes 1983) have reported that <br />piping plovers tend to nest closer to the water surface elevation than least <br />terns at sandbar colonies along the Platte River, marked differences in nest <br />elevation between species were not apparent at Missouri River colonies in 1986. <br />PRODUCTION <br />Production data for least tern breeding populations in Nebraska indicate <br />considerable variability from year to year and from location to location. <br />Along the Platte River, production rates reported at sandpit colonies in 1984 <br />and 1986 (0.42 and 0.38 fledglings /nesting pair, respectively) were twice as <br />high as reported at river sandbar colonies in 1986 (0.19 fledglings /nesting <br />pair). Moreover, of four Platte River sandbar and two sandpit colonies <br />monitored in 1982 (Ducey 1982), highest production occurred at a sandpit where <br />the owner took an active interest in protecting the nesting birds. The 30 <br />adult terns (15 breeding pairs) observed at this site produced a minimum of 10 <br />fledglings, reflecting a minimum fledging rate /nesting pair of 0.67. Within <br />the Gavins Point Dam to Ponca reach of the Missouri River, fledging <br />rates /nesting pair ranged from 0.60 to 0.16 over the 3 years of record (1982, <br />1983, and 1986), indicating nearly a four -fold difference between the highest <br />and lowest production years (1982 versus 1986). For the Missouri River reach <br />between Fort Randall Dam and Lewis and Clark Lake, production results in 1986 <br />reflected a fledging rate /nesting pair of 0.50. Production rates have not been <br />determined for breeding populations along the Niobrara or Loup rivers. <br />The low production rates reported at Platte River sandbar colonies and at <br />nesting sites along the Gavins Point Dam to Ponca reach of the Missouri River <br />in 1986 were affected primarily by flooding conditions that occurred after nest <br />establishment. Along the Platte River, flooding was the most frequent known <br />cause of direct loss of eggs during the 1986 breeding season, and losses of <br />chicks to flooding were either observed or highly suspected at most sandbar <br />colonies that year (NGPC 1986). Similarly, all Missouri River colonies showed <br />signs of water disturbance (flooding or wave action) at some time during the <br />1986 nesting season; nest scrape dampness was also evident at most sites and <br />may have contributed to low production (Schwalbach et al. 1986). <br />The overall rate of production necessary to achieve growth in the interior <br />least tern population is uncertain. However, in light of the species' <br />v <br />