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the next. Nesting habitat associated with off -river sandpits (e.g., along the <br />Platte and Loup rivers) is also transient. With the passage of time, sandpit <br />spoil piles become overgrown with vegetation or are leveled to develop housing <br />or recreation areas. Consequently, a given site may be suitable for nesting <br />one year but absent or unsuitable the next. Such changes do.not necessarily <br />represent permanent losses of habitat inasmuch as suitable habitat may be <br />forming elsewhere. As Erwin (1984) pointed out, the propensity of least terns <br />to relocate their colonies within and between years makes breeding populations <br />appear unstable, but it may be this tendency that enables the species to <br />persist despite the hazards of flooding and other disturbances. This remark is <br />equally applicable to piping plovers. <br />If suitable habitat is not available at a previously used nesting site when <br />breeding birds arrive, they may select alternate sites (Wycoff 1960) or delay <br />nesting until sandbars or beach areas become exposed (Hardy 1957). Some <br />researchers (e.g., Renaud et al. 1979) have also noted that piping plovers may <br />shift their nesting locations from year to year for unknown reasons apparently <br />not related to changing habitat 'conditions. If selected sites become <br />unsuitable during the nesting season (due to flooding, human disturbance, or <br />other factors), nest relocation or renesting may occur. Late nesting and /or <br />renesting, however, may have a lower likelihood of success because of the time <br />budget required (Schulenberg et al. 1980, Wingfield 1982). <br />The inherent ability of least terns and piping plovers to adjust to temporal <br />differences in local habitat availability is further exemplified by shifts in <br />the utilization of riverine sandbar and off -river sandpit habitat along the <br />Platte River. During recent years (1985 and 1986), sandpit spoil piles near <br />the Platte have provided habitat for nearly half the breeding birds of both <br />species that nest along this river. When flood conditions reduce the amount of <br />mid - stream sandbar habitat, sandpits near the river assume even greater <br />importance to the extent that during years of extreme and prolonged high flows <br />(as in 1983 and 1984) they provide the only nesting habitat available. These <br />findings document the fact that sand and gravel mining along the Platte River <br />has created important supplemental nesting habitat for least terns and piping <br />plovers. <br />Least terns that nest at sandpits adjacent to the Platte River forage primarily <br />in shallow areas of the river to obtain food (i.e., small fish). Foraging is <br />apparently more limited on sandpits themselves because the steeper side - slopes <br />and general lack of shallow water areas characteristic of these sites are less <br />conducive to the capture of fish prey items. Thus, even during years when <br />mid - stream sandbars are inundated during the breeding season, the river <br />continues to play an important role as a foraging area for nearby least tern <br />colonies. In contrast, piping plovers feed mainly on aquatic invertebrates and <br />can presumably satisfy their dietary requirements by foraging along shoreline <br />margins associated with either riverine or sandpit habitat. <br />COLONY SPATIAL CHARACTERISTICS <br />Available data (though limited) do not indicate greater crowding at colonies <br />during years when high flows in the Platte River restricted nesting activity to <br />off -river sandpits. Measurements of area per colony, area per nest, and <br />inter -nest distances recorded at one river colony in 1985, when moderate flows <br />prevailed during the breeding season, fell within the range of values measured <br />iv <br />