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versus 1975 is in clear contrast to Downing's (1980) prediction that, without <br />active habitat protection and enhancement, "[a]lmost complete loss of these <br />populations may occur in a few years." <br />Since the initiation of systematic surveys along Nebraska rivers, the <br />discernible trend during years of similar survey coverage and effort has been <br />one of increasing rather than decreasing populations. These increases may <br />reflect 1) the improved ability of investigators to locate nesting colonies <br />(resulting from greater experience and knowledge of the species' behavior and <br />nesting habits); 2) distributional shifts, or immigration of adult birds from <br />other nesting areas within the breeding range; 3) true population increases; or <br />4) a combination of all factors. Regardless of which factors apply, recent <br />survey results attest to the fact that habitat along these rivers can support <br />considerably larger least tern breeding populations than was previously thought <br />to be the case. <br />More recently, systematic surveys along the Platte, Missouri, and Niobrara <br />rivers have been expanded to include the collection of more complete census <br />data on piping plovers. However, because expanded survey emphasis was not <br />initiated concurrently among all rivers, insufficient data presently exists to <br />evaluate trends in the statewide population over time. Nonetheless, as in the <br />case of the interior least tern, results clearly indicate that Nebraska's <br />piping plover population is substantially larger than earlier believed, and <br />that the state is an important stronghold for this species as well. In 1986, <br />the following numbers of adult piping plovers were censused along various <br />rivers in the state: 164 along the Platte; 172 along the Missouri below Gavins <br />Point Dam; and 11 along the Missouri below Fort Randall Dam. Although the <br />Niobrara River was not surveyed in 1986, 100 birds -were censused along this <br />river in 1985. Assuming stable abundance along the Niobrara from 1985 to 1986, <br />these census results reflect a total statewide population of approximately 450 <br />adult piping plovers as of the 1986 breeding season. Since recent systematic <br />surveys have not inventoried all potential or known nesting habitat within the <br />state, this estimate should be considered conservative. <br />By way of comparison, the statewide population of approximately 450 piping <br />plovers, as determined from systematic surveys through 1986, is about nine <br />times larger than estimated in 1983 (Haig 1985) before comprehensive census <br />information was available for major breeding areas in the state. Based on a <br />review of the species' status reported by Haig (1985), the present breeding <br />population in Nebraska exceeds that of all other states in the nation, and is <br />second only to Saskatchewan among the Canadian provinces. <br />SHIFTS IN COLONY LOCATIONS <br />Interior least terns and piping plovers exhibit an opportunistic nest site <br />selection strategy that provides resiliency in responding to temporal and <br />spatial changes in the availability of suitable habitat. Systematic surveys <br />along Nebraska rivers provide clear documentation that both species make <br />frequent changes in nesting colony locations. By nature, riverine sandbars are <br />ephemeral and go through natural successional cycles of formation, shifting <br />and /or stabilization, and vegetation development. In addition, water -level <br />fluctuation, as affected by the high variability in runoff and precipitation <br />characteristic of the region, may reduce or enhance the availability of <br />riverine or interior wetland habitat either broadly or locally from one year to <br />iii <br />