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' Growth and survival rates of northern pike fry depend on temperature. <br />Survival is poor at temperatures less than 5.8° C (Lillelund 1966). Growth <br />rate is positively related to temperature for pike fry held at constant tem- <br />peratures between 7.5° C and 25.6° C, but mortality of 1-day old yolk sac fry <br />increases significantly at temperatures greater than 20.8° C; thus, the rate <br />of net biomass gain is highest at 20.8° C (Hokanson et al. 1973). The rate is <br />high for temperatures from 18.0 to 25.6° C. <br />Northern pike are spring spawners, and early <br />ence a progressively warmer average temperature. <br />tures increases within the embryo and fry stages <br />al. 1973). The optimal temperature for growth a <br />from about 21° C shortly after hatching to 26° C <br />et al. 1973). <br />developmental stages experi- <br />Tolerance of high tempera- <br />(Lillelund 1966; Hokanson et <br />nd survival of fry increases <br />after 1 or 2 weeks (Hokanson <br />Juvenile. Young of the year and yearling northern pike fed ad libitum <br />grew in weight most rapidly at 19° C; maximum growth in length occurred at <br />21° C (Casselman 1978). Growth rate increased sharply at temperatures greater <br />than 10° C but was positive (about 4°0 of the maximum) even at 3 to 4° C. No <br />growth occurred above 28° C. The growth rate- of yearling northern pike held <br />in small Missouri ponds (average depth = 1.0 m) approached zero during a 3 to <br />5 week period in midsummer when water temperatures exceeded 27° C (Weithman <br />and Anderson 1977). <br />Dissolved oxygen concentrations below 7 ppm (77% saturation) resulted in <br />reduced growth rates of juvenile northern pike at a temperature of 18.6° C and <br />a flow rate of 0.24 ml/sec (Adelman and Smith 1970b). The decline in growth <br />rate was gradual down to concentrations of 3 to 4 ppm (33 to 44% saturation) <br />and sharp at levels below 3 ppm. <br />Young of the year northern pike (12 to 38 g) showed signs of stress when <br />held for 20 to 24 hours at dissolved oxygen concentrations less than 6 ppm <br />(temperature 21 to 22° C) (Petit 1973). Feeding ceased at concentrations of <br />2 ppm, and none of the test fish survived more than 20 hours at 1 ppm._ Test <br />fish held at 1 ppm until they appeared close to death recovered within 24 <br />hours when transferred to water with a dissolved oxygen concentration of <br />6 ppm. <br />Adults. Temperature requirements of adult and juvenile northern pike <br />appear to be similar. Maximum growth in length of age 2 and 3 northern pike <br />in an Ontario lake occurred at approximately 20° C; fish of similar age reared <br />in the lab grew best at approximately 21° C (Casselman 1978). <br />Northern pike in Lake Windermere (England), where water temperatures <br />rarely exceed 21° C (Macau 1970), grew faster in warm than in cold years <br />(Frost and Kipling 1967). Surface water temperature, indexed as the cumulative <br />number of degree-days over 14° C, accounted for 86% of the variation in weight <br />of 4-year old pike, for 16 year-classes. Comparison of scale patterns for <br />northern pike from Waskesiu Lake in northern Saskatchewan with those for pike <br />from several shallow Michigan lakes (Williams 1955) suggests that high summer <br />temperatures may interrupt the growth of northern pike in more southern <br />regions. <br />11 <br />