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production can be attributed to the lateral exchange of allochthonous material and <br />nutrients between the floodplain and the main river channel during the annual flood <br />pulse. The flood pulse creates access to a variety of ephemeral habitats such as <br />backwaters, marshes, and lateral ponds. Areas of inundation are colonized by aquatic <br />biota with life histories synchronized with annual access to these favorable habitats with <br />high resource availability. The recession of flood waters transports nutrients, detritus, <br />and organisms back into the main river channel (Johnson et al. 1995). Many studies <br />provide additional evidence supporting this concept (Gosselink and Tumer 1978, Odum <br />1984, Ward 1989), especially in large, tropical riverine systems. <br />The River Continuum Concept (Vannote et al. 1980) states that rivers are <br />dependent on longitudinal linkages that transfer energy downstream. A significant <br />portion of this energy is composed of allochthonous input in the form of terrestrial <br />vegetation detritus derived from upper reach riparian and floodplain habitats. A <br />combination of the two concepts would argue that the input of allochthonous detritus <br />derived from riparian and floodplain habitats during floods is a significant source of <br />energy input to these ecosystems (Vannote et al. 1980, Junk et al. 1989, Johnson et al. <br />1995), not only for the immediate adjacent riverine system but also for downstream <br />reaches. <br />Ward and Stanford (1995) expanded on the River Continuum Concept with the <br />development of the Serial Discontinuity Concept which theorizes that dams interrupt the <br />longitudinal energy flow in river systems. Reservoirs behind dams serve to trap coarse <br />and fine particulate organic matter and interrupt sediment transport, both important to <br />longitudinal energy transfer. The rivers below these energy transfer interruptions are <br />then dependent on light or the lateral exchange of allochthonous input from floodplains <br />for energy input. The Green River below Flaming Gorge Dam is set in a similar scenario <br />of inten'upted energy flow and dependence on lateral exchanges with the floodplain for <br />energy input. <br />These lateral exchanges enhance biological productivity and maintain a diverse <br />and rich assemblage of species (Bayley 1995). Mesic conditions promote the <br />development of riparian vegetation characterized by a highly productive plant <br />community. Many invertebrates have evolved life history strategies that take advantage <br />of increased habitat availability and food resources associated with the inundation of <br />floodplain and riparian habitats (Junk et al. 1989, Goulding 1980). An invertebrate <br />assemblage characterized by increased density and species diversity develops in <br />association with cyclic floodplain inundation (Sparks 1995, Bayley 1995, Allan and <br />Flecker 1993). <br />River ecosystem productivity has been shown to be limited by nitrogen and <br />phosphorous availability in some instances (Newbold et al. 1983, Tiessen et at. 1994). <br />In large river systems, significant available portions of the these nutrients are derived <br />from floodplain habitats. Floodplains influence food web productivity through the input <br />of limiting nutrients from the bottom up in a hierarchical manner (Newbold et al. 1983, <br />Vought et al. 1994, Leonardson et al. 1994). Nutrients in solution, derived from <br />floodplain detritus and soils, are available for autochthonous production <br />(photosynthesis) in the growth of phytoplankton and periphyton. This process initiates <br />the assimilation of floodplain nutrients and potential productivity into aquatic food webs. <br />1.3 <br />