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MAYDEN ET AL.-SYSTEMATICS OF WESTERN CYPRINIDS <br />12 <br />10 <br />L <br />.O <br />L <br />O <br />L <br />O <br />C <br />'b <br />. b <br />b <br />' b <br />b <br />0 <br />Y as 16 • ? bb ' o <br />?a a, <br />?"bsb <br />ad'O/. bp?b p ?' ? .' <br />Qa 0 <br />o o <br />9, <br />I-a0a?9? i <br />0 <br />21- <br />40 <br />60 80 <br />100 120 <br />Standard Length <br />829 <br />Fig. 4. Linear regression of interorbital width (vertical axis) with standard length (horizontal axis). The <br />"o" represents P. oregonensis and the "a" and "b" represent P. umpquae collections UCLA W81-15 and UCLA <br />W86-4, respectively. The dotted lines indicate the area included in the 95% sample prediction interval for <br />each regression line. <br />Mylopharodon conocephalus to the genus Plycho- <br />cheilus and a nested series of sister-group rela- <br />tionships between P. lucius and A grandis, P. <br />umpquae, and P. oregonensis. The PAU P analysis <br />using all unordered characters yielded a single <br />tree of identical topology with a CI of 0.943 <br />(with homoplasy masked). <br />Hesperoieucus symmefricus <br />DISCUSSION <br />That all four altopatric species of Plychoched us <br />can be distinguished morphologically is sur- <br />prising only in regard to the traditionally en- <br />countered difficulty in identification of P. <br />umpquae and P. oregonensis from single charac- <br />Myiopharodon conocepha/us <br />I L P. grondis <br />P. umpquoe <br />P. oregonensis <br />P iucius <br />Fig. 5. Wagner tree of relationships among species of Mylopharodon and Ptychocheilus. Hesperoleucus sym- <br />metricus is the outgroup. Limb length is in proportion to the number of character changes. The consistency <br />index = 0.868.