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830 COPEIA, 1991, NO. 3 TABLE 5. GAP-CODED TAXON X CHARACTER MATRIX. Character numbers are the same as in Table 1. Ptycho cheilus umpquae is represented only by specimens from the Umpqua drainage. Character codings 1-10 11-20 21-30 31-40 41-51 H.symmetricus 0021002000 2033010011 0011112203 0231101212 03300010000 M.conocephalus 0010002000 1022010011 0011011112 1120101121 02201100111 P.lucius 0000000000 0000000000 0000000020 2000000030 10023211010 IG P.grandis 1010001000 1011010000 0011000202 0010001101 01023100110 P. oregonensis 1010001000 1021010011 0011000201 0010001101 01112110110 P.umpquae 1010001000 0011010011 0011000202 0010001101 01122110110 to ters alone. Although P. umpquae is known to interpretation supplied by C. Bond (pers. comm. have "smaller scales" along the predorsal mid- to D. Buth, 27 April 1982) was especially per- line and above the lateral line than P. oregonensis tinent: (Snyder, 1908; Eddy and Underhill, 1969; Car- The Umpqua population was derived C ney and Page, 1990), these counts are difficult. re lar th diff l d l from the Coast Fork of the Willamette Riv- e gu erences, e row orsa For pre sca er probably near the two drainage pass at d 1, scale rows illustrated in Snyder (1908) reflect Divide, whereas the Siuslaw population was h1 considerable artistic license. The scales actually derived from the Long Tom River, a Wil- form extremely irregular rows, making it dif- tributary, via stream capture near lamette ficult to obtain reliable counts. Mean values of Goldson. I think the two populations are lateral-line scales differ between these species related only through their old Willamette (Table 2), but overlap for these counts is suffi- River cousins and have converged on the cient to prevent the character from distinguish- character of small scales. Between the Sius- ing all members of these two forms. The gap- law and the Umpqua are two separate coding procedure used herein recognized the drainages-Siltcoos and Tahkenitch-that same state for lateral-line scales for P. umpquae had squawfish in Evermann and Meek's j and P. oregonensis. Both species may be reliably s of transverse scale b i i h d b di time, naturally or otherwise. An old timer y num er ngu s e st T bl me the re- at Tahkenitch Lake showed a e rows and circumpeduncular scale rows ( mains of a "bait boat" that was used on the 8)' Siuslaw in the 1890's (and before?). Bait Bond (pers. comm.) has expressed concern fishermen ran the Siuslaw in the long, nar- that any difference recognized between all pop- row craft and caught "minnows" that they ulations of P. umpquae and P. oregonensis may be sold for trout fishing (later, bass fishing) in ,I' simply a result of the greater scrutiny to which the lakes. 1 have wondered if the bait in- these local populations have been subjected and dustry was not responsible for the spread that comparable variation may exist among oth- of some of the Siuslaw fishes. Squawfish are er populations of P. oregonensis. Although we not numerous in the lakes now. Woahink have not studied P. oregonensis in detail through- out its range, our samples from localities as dis- tant as western Oregon and Montana showed TABLE 6. HOMOPLASY MATRIX FOR THE CLADOGRAM some differences but nothing so pronounced as (FIG. 4). is seen between all populations of this species examined and P. umpquae. However, if such is Taxa 1 2 3 4 5 6 the case and diagnosable forms do exist within • H. symmecricus 0 P. oregonensis, then this diversity should be rec- 2. M. conocephalus 0 0 . ognized because it may have significant conse- 3. P. ducius 6 12 0 quences for the preservation of forms in need 4. P. grandis 2 4 0 0 rill rotection. Of protection. P. oregonensis 0 0 2 2 0 5. l+" The observation that P. umpquae may com- i b . P. umpquae 6 e sur r s- rise more than one form may not prise Y P (Umpqua drainage) 0 0 0 0 0 0 li ?i i? ing. The following background information and