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<br />222 <br /> <br />GREAT BASIN NATURALIST <br /> <br />[Volume 55 <br /> <br />11m2 on over-flooded meadows in Czechoslo- <br />vakia, In Lake Norman, NC, Bowen (1983) <br />reported a mean larval ceratopogonid density <br />of 767/m2. <br />Ceratopogonid densities reached a peak in <br />the August river channel sample (13,0261 m2)_ <br />much higher than any reported in the litera- <br />ture above. In their study of the Green River, <br />Grabowski and Hiebert (1989) did not report <br />densities, but did conclude that ceratopogo- <br />nids were more abundant in river channel <br />samples than in backwaters. Our study supports <br />this conclusion, Average densities for the river <br />channel July and August samples were 3608/m2 <br />and 13,026/m2, respectively, compared to 961m2 <br />and 461/m2 for the backwater July and August <br />samples. Ceratopogonid larvae were complete- <br />ly absent from the ephemeral side channel as <br />well as the June and July seasonally inundated <br />wetland samples. <br /> <br />Chironomidae <br /> <br />Chironomidae are typically the most abun- <br />dant macroinvertebrates in lentic (Strayer 1985) <br />and lotic (Grzybkowska and Witczak 1990) sys- <br />tems. Studies of relatively small geographical <br />areas have reported impressive numbers of <br />species. For instance, Douglas and Murray <br />(1980) found 142 species in Killarney Valley, <br />Ireland. High diversity of chironomids makes <br />them important as indicators of environmental <br />condition (Wingard and Olive 1989). They are <br />also abundant and provide an important food <br />source for fish (Brown et al. 1980, Winkel and <br />Davids 1987, Grabowski and Hiebert 1989), <br />waterfowl (Titmus and Baddock 1980), and <br />other migratory birds (Bowman 1980). <br />We identified 19 chironomid genera from <br />our sites within the Green River ecosystem. <br />Other investigations oflotic systems have yield- <br />ed similar numbers-12 genera in the upper <br />Tuscarawas River, OH (Wingard and Olive <br />1989), 24 genera in the River Frome, England <br />(Pinder 1980), 25 genera in the Mississippi <br />River (Beckett et al. 1983), and 36 genera in <br />Juday Creek, IN (Berg and HellenthaI1991). <br />Grabowski and Hiebert (1989) studied the <br />Green River in the same general area consid- <br />ered in our study and also identified 19 genera. <br />However, only seven of the genera reported <br />by the latter authors were found in our study: <br />Chironomus, Cricotopus, Cryptochironomus, <br />Polypedilum, Procladius, Tanypus, and <br />Tanytarsus. <br /> <br />Densities of chironomids in aquatic sys- <br />tems can vary substantially. In a study of Lake <br />Vissavesi, Finland, Paasivirta and Koskenniemi <br />(1980) reported densities of 641m2 in a coarse <br />debris habitat and 2997/m2 in a moss-grown <br />site. Jonasson and Lindegaard (1979) reported <br />59,000/m2 from Lake Myvatn, Iceland. Vari- <br />ability in lotic systems has also been docu- <br />mented. Pinder (1980) reported densities from <br />a low of 481m2 to 6273/m2 in a chalk stream in <br />England, and Grzybkowska (1989) found <br />1O,664/m2 in the River Grabia, Poland. While <br />no distinct trends exist when comparing chiro- <br />nomid densities in lentic and lotic systems, den- <br />sities are influenced by sediment size (Paasivirta <br />and Koskenniemi 1980, Beckett et al. 1983). <br />Chironomid densities from the July and <br />August river channel samples were 4148/m2 <br />and 3516/m2, respectively. River backwater <br />samples were 31,125/m2 and 22,864/m2 for the <br />same times. Grabowski and Hiebert (1989) <br />reported maximum chironomid densities in <br />the same area of the Green River of less than <br />100/m2 for the river channel and 2800/m2 for <br />river backwaters-substantially less than our <br />estimates. It is possible that annual differences <br />in seasonal discharge, area of the sampling <br />device, and later sampling period all contrib- <br />uted to this discrepancy. However, because of <br />significant differences in mesh size (63-tLm <br />ours, 600-tLm Grabowski and Hiebert's), data <br />of Grabowski and Hiebert and ours cannot be <br />considered equivalent. It is worth noting that <br />mesh sizes larger than 100 tLm have been shown <br />to negatively bias density estimates (Strayer <br />1985). <br /> <br />Community Similarity <br /> <br />Cluster analysis of the data showed that, in <br />general, habitat types clustered together inde- <br />pendent of sample date, suggesting that the <br />different habitat types studied in the Green <br />River are distinct. Beckett et al. (1983), for ex- <br />ample, studied five habitats in the Mississippi <br />River and also found them to remain composi- <br />tionally distinct regardless of flow and sample <br />date. Distribution and abundance of benthic <br />macroinvertebrates characteristic of these <br />habitat types have been attributed to flow con- <br />ditions and sediment size in our study. Since <br />flow conditions are the major determinant of <br />particle size, flow conditions are likely the <br />determining factor. This conclusion has also <br /> <br />t <br />