<br />1995]
<br />
<br />MACROINVERTEBRATES OF THE GREEN RIVER
<br />
<br />223
<br />
<br />! (
<br />
<br />been reached by other investigators (Beckett
<br />et al. 1983, Statzner and Higler 1986).
<br />Grabowski and Hiebert (1989) concluded
<br />that benthic macroinvertebrate densities in
<br />backwaters of the Green River were higher than
<br />those of the river channel. Our data suggest
<br />that the seasonally inundated wetland and
<br />ephemeral side channel are also valuable habi-
<br />tats and have the potential to contribute sub-
<br />stantial biomass to the Green River system.
<br />Oligochaete and chironomid densities report-
<br />ed in our study are comparable to other lotic
<br />systems (Koehn and Frank 1980, Pinder 1980,
<br />Grzybkowska 1989, Grzybkowska and Witczak
<br />1990, Palmer 1990). High densities of nema-
<br />todes and ceratopogonids imply that these
<br />groups may be very important in the overall
<br />energetics of the Green River system, Both
<br />should be studied more intensely. The overall
<br />dynamics of these communities is undoubtedly
<br />associated with seasonal changes in flow as well
<br />as year-to-year variability in annual discharge.
<br />This study, while describing a backwater, river
<br />site, side channel, and floodplain wetland over
<br />a short time interval, does not allow a full
<br />assessment of either annual or spatial variabil-
<br />ity, It is clear that some sort of successional
<br />colonization of various habitats occurs; for
<br />instance, floodplain wetlands are maximum in
<br />extent during highest spring-early summer
<br />flows, but their faunal development lags peak
<br />flooding. Backwaters do not exist during high
<br />flows, but as floodplains diminish with reced-
<br />ing water levels, backwater habitats develop.
<br />Again their faunal assemblages tend to lag be-
<br />hind the emergence of recognized backwaters.
<br />While we documented what appears to be
<br />seasonal succession within habitat type, such
<br />changes should not be assumed the norm.
<br />Until a detailed study is undertaken for the
<br />Green River or Colorado River system with
<br />replicate habitats over at least a full year period,
<br />our observations must be considered tentative,
<br />Further, annual discharge can vary tremen-
<br />dously from year to year, depending upon fac-
<br />tors such as drought cycles and their link with
<br />EI Nino dynamics in the Pacific. Thus, what is
<br />seen in one year may not be representative of
<br />all years. Such factors introduce additional
<br />variables that should be considered when
<br />attempting to understand the dynamics of the
<br />benthos of the Green River.
<br />
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