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backwaters and embayments are used by adult Colorado squaw- <br />fish and razorback sucker as refuges and staging areas during <br />runoff and as foraging areas during low flow. The require- <br />ments of these target species cannot be universally satisfied <br />with construction of a single type of feature. Enhancement <br />of high-water and low-water habitats is needed. <br />Interesting comparisons can be made between natural and <br />manmade backwaters and embayments in the Yampa and Upper <br />Colorado Rivers that reveal some of the more desirable <br />aspects of these habitats. Diked side channels, built for <br />erosion control, appear to offer valuable fish habitat during <br />high flows, when the fish tend to leave the main channel and <br />seek refuge in quiet areas. Adult Colorado squawfish use <br />artificial backwaters as they use naturally-occurring <br />ones. Diking side channels in areas inhabited by adult <br />squawfish and razorback sucker warrants consideration for <br />recreating backwaters lost to reduce flows and channeliza- <br />tion. Earthen dikes placed at upstream ends of side channels <br />to prevent the river from flowing through allow flooding from <br />the lower end and are a desirable manmade structure. Dikes <br />should not be allowed to breach, and side channels should <br />have gentle gradients that allow backwaters to drain with <br />descending flows and prevent the establishment of potentially <br />competitive non-native fish populations. <br />Habitat for juvenile and yoy squawfish and razorback <br />sucker can be created from partially-breached dikes at heads <br />of side channels. This allows the channels to flush silts <br />and particulates during runoff, but to become slow-flowing <br />backwaters during low flow that support high levels of <br />macrophytes and insects for young fishes. Care .should <br />be taken before creating these low-water habitats for native <br />species since the presence of high densities of non-native <br />species may pose serious competition/predation problems. <br />Embayments can also be created to provide low-flow nur- <br />series for young native fishes by diverting the river energy <br />into an eddy effect. Wing dams and reinforced banks can cre- <br />ate this digging action, but more study is needed to under- <br />stand what substrates respond best to various strucCures. <br />Abandoned gravel pits constitute unnatural riverine <br />features that allow competitive and predaceous non-native <br />fishes to flourish because they remain sheltered from- the <br />river year around. These large, unnatural features should be <br />permanently sealed from tfie river and managed as warm-water <br />sport fisheries. Subterranean flow into these pits is <br />sufficient to maintain good water levels and quality. <br />Gently-graded gravel pits that flood during runoff but drain <br />534 <br /> <br />with descending flow may benefit native fishes by providing r <br />staging or resting areas during runoff while precluding - <br />permanent populations of non-native fishes. <br />Habitat enhancement to recover native fishes should <br />utilize natural riverine features, where possible. Existing <br />side channels, tributaries, old river channels and oxbows <br />should be utilized for backwaters to insure a stable, long- <br />lasting feature. The feasibility and expense of diking these <br />channels will depend largely on access and side effects of <br />the procedure. Excavating new channels or backwaters is not <br />advised because the shifting na-Cure of the river is likely to <br />fill these with sand and silt and render the operation <br />a wasted expense. <br />SUMMARY <br />1. Backwaters created by diked side channels resemble in <br />appearance and function natural habitats. These benefit <br />protected native fishes by providing resting, staging <br />and feeding areas during high flow, and drain at low <br />flow to prevent establishment of permanent populations <br />of competitive non-native fishes. <br />2. Backwaters that persist at low flow are used by native <br />fishes for foraging and resting. These are generally <br />inundated and flushed of non-native fishes at high <br />flows. <br />3. Side channels with breached dikes can provide low- <br />velocity nursery habitats that flush of silts at high <br />flow. These features resemble naturally-occurring <br />habitats that can harbor large numbers of non-natives; <br />so, community structure must be well known before <br />constructing breach able dikes on side channels. <br />4. Embayments are important low-water nurseries and habi- <br />tats for young fishes. These can be created with wing <br />dams and reinforced river banks to direct river energy <br />into an eddy effect that digs the embayment and builds <br />protective sand bars. Embayments are flushed of silt by <br />annual runoff and their short-term nature lessens <br />potentially competitive effects of non-native species. <br />5. Large, deep, open-water gravel pits that are refuges for <br />non-native fishes but attract native fishes during <br />runoff should be sealed from the river and managed as <br />warm-water fisheries and wildlife sanctuaries. This may <br />reduce problems for native fishes associated with <br />competition, predation and pathogens, and reduce the <br />influx of non-native fishes into the river system. <br />SAS <br />