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main channel because larger fish are less susse tible to predation by nonnative fish. These <br />potential benefits could be offset, how er, if no ative fish species also experience enhanced <br />growth and survivorship. In fact, so AW ive sp 'es may gain advantage because they are <br />adapted to the lentil environment of the p ain wetland. Therefore, management and <br />maintenance of these wetlands require considerable understanding of the community ecology of <br />native and nonnative species and their interactions. <br />Additionally, the enhanced productivity of floodplain wetlands may be important in enhancing <br />survivorship of adult native fish. Modde (personal communication), Osmundson and Kaeding <br />(1989), Tyus and Karp (1990, 1991), and Holden and Crist documented the use of floodplain <br />wetlands or low velocity areas by adult razorback sucker. Wick (personal communication) <br />speculated that adult fish use highly productive areas to replace valuable energy reserves <br />exhausted while spawning. Resting habitats, therefore, may be important to enhance the <br />survivorship of the adult population of razorback suckers in the Upper Basin. <br />Levee Removal Issues <br />The following paragraphs list primary issues concerning the floodplain habitat enhancement <br />component of UCRBRIP. We have listed each issue either as a statement or a question. Each <br />issue is followed by a brief discussion. The discussion of each issue is in the levee removal <br />strategy. <br />Naturallv Flooding Wetlands will be Used to Develop Management Criteria. To date, most of <br />the effort on the benefits of floodplain function to endangered species within UBRIP has focused <br />on "manipulated" sites that use human intervention to control flooding. A substantial amount of <br />area floods naturally in the Colorado River system, yet little, if any attention, has been given to <br />naturally flooding wetlands in the basin even though it appears that a substantial amount of <br />flooding occurs in some areas during high flow years (e.g., 1983, 1984, 1995; Crowl, personal <br />observation). This strategy added to information from previous studies allows managers to <br />determine fish production dynamics in a range of natural floodplain wetlands and specific <br />restorations (levee removal) sites as they become available. This document outlines a temporal <br />and spatial strategy to compare fish population response and dynamics of 'restored' to that of <br />natural floodplain areas. This plan contains a monitoring protocol designed to determine fish food <br />(zooplankton, macroinvertebrate, etc.) and fish population responses (both native and nonnative) <br />associated with these wetlands. <br />Role of Manipulated Sites. Manipulated sites (i.e., Old Charlie, Walter/Walker, Johnson, Boat <br />Bottom, etc.) will be used to address specific questions concerning floodplain function. Sites with <br />control structures provide an opportunity to answer questions that cannot be addressed in existing <br />natural wetlands, particularly as they relate to timing and duration of inundation. In addition, they <br />also offer an opportunity to use the control structures to increase the effectiveness of fish <br />sampling. They also allow the opportunity to determine the effectiveness of different gears in <br />5