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Introduction <br />Inundated floodplains and terraces, hereafter referred to as floodplain wetlands, constitute a <br />highly productive component of the riverine ecosystem. In the Colorado River system, floodplain <br />wetlands consist of shallow, slow-moving water typically with higher nutrient concentrations, <br />primary productivity and invertebrate densities than adjacent areas of main channel flow (ongoing <br />studies by Shiozawa; Cooper and Severn 1994 a, b, c, and d; see Forsberg et al. 1993). These <br />wetlands may serve as essential rearing and feeding areas for some native fish species, including <br />the endangered razorback sucker (Xyrauchen texwms) and Colorado squawfish (Ptychocheilus <br />Lucius). The Upper Colorado River Basin Recovery Implementation Program (UCRBRIP) has <br />targeted acquisition and reinstitution of active floodplain wetlands (within the 100-year <br />floodplain) to main channel flows as a priority recovery activity. However, effective management <br />of these wetlands for recovery of endangered fish requires considerable understanding of <br />Colorado River wetland ecology. The necessary information will be acquired through research <br />and experimentation which is implemented on an increasing temporal and spatial scale. This <br />schedule allows evaluation of preliminary efforts and subsequent modification to prevent <br />prodigious effort and expense at the onset of the project. <br />Background <br />Previous research has documented the importance of main channel low velocity areas (e.g., <br />backwaters) as native fish habitat (Tyus). More recent research has focused on the <br />geomorphology of nursery habitats (Pucherelli; Rakowski and Schmidt) and their ecological <br />function in Colorado squawfish (Lentsch) and razorback sucker population dynamics (Vernal, <br />CRFP). Floodplain wetlands may be equally important as biologically complex habitat for <br />endangered fish. Furthermore, a reduction of channel and floodplain complexity associated with <br />dam construction, flood control and riparian vegetational changes over the last 100 years <br />(Andrews 1986) lends urgency to understanding and restoring a functional, integrated floodplain <br />system in the Upper Colorado River system. <br />Floodplain wetlands are generally areas of high primary productivity due to elevated water <br />i temperatures, nutrient levels, and light intensities. Decaying plant material (emergent and flooded <br />terrestrial vegetation) supports a large portion of the decompositional pathway within the riverine <br />foodweb. Consequently, macroinvertebrate production is high because of an enriched food base <br />(phytoplankton, etc.), a structurally complex environment (structure provided by emergent plants, <br />flooded vegetation, and algal mats), and relatively warm water temperatures. These factors taken <br />in aggregate suggest that floodplain wetlands are extremely productive habitats. <br />Because of enhanced productivity of food organisms, floodplain wetlands facilitate rapid fish <br />growth within the floodplain (Osmundson) and survival within the main channel. Growth of <br />native young-of-the-year (YOY) fish in floodplain wetlands is enhanced because of greater food <br />abundance and higher growth rates; subsequently, survivorship of YOY fish is enhanced in the <br />4