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newly created habitat, and for zooplankton blooms to occur (see Cooper and Severn <br />1994b). A minimum duration of inundation that assures time for these two processes to <br />occur as well as provide time for larvae to feed and grow may be 3-4 weeks. <br />Within-channel Productivity <br />Assuming that reproduction and survival of young can be enhanced for Colorado <br />River populations of razorback sucker and Colorado pikeminnow, carrying capacity of their <br />environment will become the next factor that limits population size. Providing the <br />maximum amount of preferred mesohabitats partially addresses this constraint. In addition <br />to limitations imposed by physical habitat are the limitations of food availability. Habitats <br />containing abundant food will support more fish than those that do not. To provide more <br />food for the endangered fish, primary and secondary productivity should be maximized. <br />Algae and detritus form the base of the riverine food web, directly supporting invertebrates <br />and some fish. Adult razorback suckers feed on benthic and drifting invertebrates, algae, <br />and detritus (Bestgen 1990), as do sympatric bluehead suckers Catostomus discobolus and <br />flannelmouth suckers C. latipinnis (Osmundson 1999). Periphyton and terrestrial inputs of <br />organic debris are the source of detritus. Most terrestrial organic debris enters the river <br />during spring when high flows flood the banks and entrain shoreline and bottomland <br />accumulations of branches and leaf litter. For periphyton production, clean rock surfaces <br />for attachment sites are required as well as sufficient water clarity to allow light penetration <br />to the river bed. Invertebrates feed on algae attached to rock surfaces and on detritus both <br />in the drift and in the interstitial spaces among coarse substrate particles. In addition to <br />food, invertebrates, like fish, require certain physical habitats: these include rock surfaces <br />for attachment sites and interstitial spaces for shelter (Waters 1995). Fish that subsist on <br />algae, detritus and invertebrates in turn provide forage for the piscivorous Colorado <br />pikeminnow (Osmundson et al. 1998, Osmundson 1999). <br />To promote within-channel productivity, high spring flows are needed to clean <br />gravel-cobble substrates (Osmundson et al. 2001). In the absence of flows of sufficient <br />magnitude fine sediment (silt and sand) accumulates in the river bed filling the spaces <br />28