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required by many invertebrate species (Osmundson and Scheer 1998, Osmundson et al. <br />2001). If the tops of rocks become covered with fine sediment, algal production may also <br />decline. To flush fine sediment from the bed, flows must be high enough to dislodge and <br />move the larger particles. Gravel river beds are typically composed of surface and <br />subsurface layers. The surface layer, sometimes called a pavement or mobile armour, is <br />coarser than the subsurface material. Milhous (1973) found that mobile armour plays an <br />important role in the deposition and retention of fine sediment. At low flows, the immobile <br />surface layer acts as a sink for suspended sediment, which deposits at the interface between <br />the pavement bottom and the subsurface top. This zone acts as a silt reservoir: at high <br />discharge the pavement is set in motion; the bed then becomes a source of suspended <br />sediment as fines are winnowed out. Milhous (1973), O'Brien (1987) and Wilcock et al. <br />(1996) have emphasized the necessity of surface layer mobilization for removing fines <br />below the surface layer. Pitlick and Van Steeter (1998) found that the minimum flow <br />necessary to produce widespread movement of the bed in the 15-mile reach, and the two <br />reaches immediately downstream of the Gunnison River inflow (18-mile and Ruby- <br />Horsethief Canyon), corresponded with the bankfull flow. As discussed above, the <br />bankfull flow in De Beque Canyon is approximately 20,500 cfs and in the De Beque-to- <br />Rifle reach, approximately 22,000 cfs. In the 15-mile reach, the bankfull flow is <br />approximately 21,500 cfs (Pitlick and Cress 2000). <br />Thus, bankfull flows not only provide razorback sucker larvae and adults with <br />critically important off-channel habitats and entrain organic debris from bottomlands into <br />the main channel, they also serve to mobilize within-channel substrates, flushing fines from <br />the bed and optimizing living space for macroinvertebrates. Ideally, these high spring flows <br />should then be followed by base flows with low turbidity to promote algal production. <br />Colorado Pikeminnow Spawning Cues <br />Cues that trigger Colorado pikeminnow to undergo spawning migrations and to <br />initiate spawning appear to be largely controlled by river flows and temperature. Adults <br />begin migrating to spawning areas as peak runoff declines and water temperature increases <br />29