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Hiebert 1989, Mabey and Shiozawa 1993), underscoring the importance of larval access to <br />floodplain habitats. <br />To appreciably flood bottomlands along the upper Colorado River, flows must <br />exceed the bankfull level. Pitlick and Cress (2000) estimate the median bankfull discharge <br />in De Beque Canyon (upstream of the upper diversion structure) is approximately 20,500 <br />cfs (based on field measurements of bankfull characteristics of eight evenly spaced cross- .. <br />sections). In the alluvial reach upstream (De Beque-to-Rifle), where adjacent bottomlands <br />are present, the estimated median bankfull discharge is 22,000 cfs (based on 24 cross <br />sections). <br />In the Carter et al. (1985) mapping study, those habitats that exhibited predictable <br />and regular changes with flow were generally associated with physical features of the river <br />that were inundated when the river flowed beyond its normal channel banks (backwaters, <br />flooded woodlands and rubble flats) or were manifestations of river hydraulics at various <br />stages (rapids, runs and eddies increased with discharge; riffles decreased). Total area of <br />backwaters declined when flows increased from 1,710 to 9,000 cfs, but then increased 50- <br />fold at discharges over 10,000 cfs. Some flooded bottomlands were present at discharges <br />of 10,000-21,000 cfs, but were nonexistent at flows less than 10,000 cfs. As discharge <br />increased above 21,000 cfs, total area of flooded bottomlands began to increase <br />exponentially, indicating a threshold for significant over-bank flooding somewhere between <br />21,000 and 23,400 cfs. This inflection point corresponds with the median bankfull <br />discharge of 22,000 cfs calculated by Pitlick and Cress (2000) for the entire De Beque-to- <br />Rifle reach. <br />Successful year classes of razorback sucker largely depend on larvae being placed in <br />habitats containing adequate densities of forage; such densities are largely restricted to <br />flooded bottomlands. The necessary duration of flooding may depend on the time required <br />for larvae to feed during their critical first phase of life. The timing, density, size, and <br />duration of zooplankton availability must `match' the timing of the swim-up stage of fish <br />larvae. Razorback sucker larvae must find food of the right size and density within 8-19 <br />days of swim-up or they will exceed the point of irreversible starvation (Wydoski and Wick <br />1998). After inundation, some time is required for larvae to drift into or hatch within the <br />27