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by low stable base flows. The cause-effect explanation offered by McAda and Ryel (1999) <br />is as follows: high, bed-mobilizing flows are needed to create and maintain ideal substrate <br />conditions for egg deposition and incubation and the effect of this sorting and cleaning of <br />the gravel-cobble substrate is carried over into the following year when moderately high <br />flows are then adequate to remove any additional fine sediment deposited between runoff <br />events; moderate flows also allow for earlier spawning, a longer first-year growing season, <br />and more backwater habitats for the small fish to settle in. In conclusion, for high YOY <br />numbers, two conditions need to be met: high larval production in summer followed by high <br />larval survival and retention in the nursery area until fall. This combination occurs when a <br />particular set of hydrological conditions are provided. <br />Diet, growth, body condition and carrying capacity <br />Once Colorado pikeminnow are over a year old they switch from a diet consisting of <br />invertebrates to one consisting almost entirely of fish (Vanicek and Kramer 1969, Muth and <br />Snyder 1995). Backwaters are good nursery habitats because they are warm, lack current, <br />and contain relatively abundant supplies of zooplankton, macroinvertebrates, and small- <br />bodied fish (Graboswski and lEebert 1989). Even after Colorado pikeminnow move to <br />main-channel habitats, small-bodied fish, primarily non-native minnows, provide a <br />substantial part of their diet until pikeminnow reach a length of at least 550 mm. Larger <br />individuals are thought to require larger forage items (Osmundson et al. 1998), as is the <br />case with other warm-water piscivores such as northern pike Esox lucius and muskellunge <br />E. masquinongy (Scott and Crossman 1973, Gillen et al. 1981, Diana 1987). Information <br />from limited stomach content data suggests that relatively large, soft-rayed, fusiform-shaped <br />fish are eaten, including various species of sucker. Consumed suckers were as long as 47% <br />of the Colorado pikeminnow's length (summarized by Osmundson et al. 1998). <br />Theoretically, positioning of Colorado pikeminnow within the river, at both macro- <br />and micro- scales, is driven primarily by growth maximization, and growth is largely <br />dependent on the interaction between temperature and food availability (Weatherley 1972). <br />Achieving maximum growth enhances the ability of the individual to survive and reproduce. <br />When food availability is low, growth slows and body condition declines. To the extent <br />18