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possible, Colorado pikeminnow may prevent this by selecting foraging sites with high rates <br />of energy return (food) for energy expended (foraging activity). Good conditions for <br />growth include suitable temperatures in combination with relatively high availability of <br />forage fish. The benefits of high forage density may be enhanced by a combination of <br />physical habitats and river features that facilitate efficient foraging, thereby promoting fish <br />growth and allowing more Colorado pikeminnow to occupy a given reach of river (i.e., <br />forage availability as opposed to forage abundance). Supportive evidence for this includes: <br />(1) the dispersal of adults in the Colorado River to reaches upstream of Westwater Canyon <br />where native forage is most abundant (Osmundson et al. 1998), (2) the relatively high <br />density of adult Colorado pikeminnow in the 18-mile reach (downstream from the Gunnison <br />River confluence) where total area of specialized habitats (non-run habitats) is highest <br />(Osmundson et al. 2001), (3) the preference for river segments that contain a complex of <br />habitat types, as opposed to simple, single-thread, run-dominated segments (Osmundson <br />and Kaeding 1991), and (4) the preference for certain habitat types, such as pools, eddies, <br />etc. (previously discussed). Because habitat and food are so tightly interrelated, it is <br />difficult to separate selection for food from selection for habitats that allow efficient <br />foraging (Magnuson et al. 1979). Nevertheless, to promote growth of individual Colorado <br />pikeminnow and maximize carrying capacity of the river, abundant forage and a diversity of <br />habitats are important. <br />SEASONAL PARTITIONING OF THE YEAR <br />To provide favorable habitat for the endangered fish, flows must change in a <br />seasonal manner corresponding to season-specific habitat needs. Osmundson et al. (1995) <br />blocked months into seasons by analyzing habitat-use patterns of each species and <br />identifying changes in behavior that marked the beginning or end of seasons. Though <br />coexisting under the same conditions, the different behavioral patterns of razorback sucker <br />and Colorado pikeminnow result in a year that is partitioned somewhat differently (Fig. 3). <br />However, because only one flow regime can be recommended for the river, a third seasonal <br />19