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<br />I <br />I <br />I <br />I <br />I <br />I <br />1 <br />I <br />1 <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />thicker in reach 3, but backwaters often had multiple ice layers indicating a greater frequency of ice <br />jams. Ice lensing was riot observed in reach 3 because of the lack of flow fluctuation and the more <br />constant water temperature. <br /> <br />Average free water depth in backwaters varied from 0.7 to 2.9 feet in year 1 and from 0.7 to 3.2 feet <br />in year 2. Although ice formed solid to the substrate in shallow areas (<0.5 feet) of most backwaters, <br />we did not observed ice development extensive enough to substantially reduce free water in any <br />backwater. <br /> <br />Average water temperature in backwaters varied from 0.06 to 3.660 C in year 1 and from 0.10 to <br />4.780 C in year 2. Despite measurement of supercooled water temperatures in the mainstem (<00 C), <br />supercooled conditions were not recorded in backwaters during this study . We believe that frazil ice <br />development was not extensive enough to jam backwaters beneath surface ice and create the <br />supercooled condition. <br /> <br />Average dissolved oxygen varied from 6.0 to 13.6 mgll in year 1 and from 5.1 to 16.3 mgll in year <br />2. Most recorded concentrations were at or just below saturation for the given temperature and <br />elevation. Low concentrations of 5.1 and 5.3 mgll in two backwaters in year 2 were attributed to a <br />local biological oxygen demand from a dense layer of dead leaves and vegetation, as well as fecal <br />pellets, associated with a beaver pond. The high concentration of 16.3 mgll was recorded in a <br />backwater that was isolated from the main channel by an ice plug formed at the mouth. A substantial <br />amount of periphyton was present on the cobble substrate and the ice was clear and absent of snow <br />cover, suggesting active photosynthesis and oxygen production in the closed system. <br /> <br />In summary, of the 23 backwaters observed and measured in the winters of 1993-94 and 1994-95, <br />the only factors that made these nursery habitats unsuitable for young fish were natural geomorphic <br />changes and ice jams that caused flow through the backwaters and negated their value as sheltered, <br />low-velocity habitats (Tables 9, 10). While these changes were caused indirectly by ice conditons, <br />no detrimental conditions were seen in backWaters as a direct result of ice conditons. Those <br />backwaters that persisted retained greater than the minimum area of30 m2, water depth of9.0 em <br />or greater, water temperature remained above -0.50C, and dissolved oxygen remained above 5.0 <br />mgIl. <br /> <br />7.0 DISCUSSION <br /> <br />Winter provides one of the most stable environments for riverine fishes of any season of the year <br />(Hynes 1969). Natural flows are relatively stable and an ice cap often dampens fluctuations (Wick <br />and Hawkins 1989) and maintains above-freezing, isothermal conditions. Lowered water <br />temperatures reduce community metabolism and fish respiration, sending fish into a state of reduced <br />activity (Schmidt et al. 1987). <br /> <br />Since Flaming Gorge Dam was built on the Green River in 1963, winter flows are higher by an <br />average of over 1,000 cfs for each month. Hypolimnetic releases of 8-140 C in summer have kept the <br />river relatively colder, and releases of 4-80C in winter have kept the river relatively warmer than pre- <br />dam conditions. These thermal effects are greatly ameliorated by distance and by the Yampa River, <br /> <br />20 <br />