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<br />Table 3. Mean number of aquatic organisms per liter (combined species of <br />zooplankton and free-swimming benthic invertebrates) collected <br />in five vertical tows with a 20.3 cm plankton net from three <br />backwaters in the lower Green River during 1997. ~/ <br /> <br />I <br />I <br /> <br />I <br /> <br />Date Millard Canyon Anderson Canyon Holeman Canyon <br />May 1 6.2 2.4 6.6 <br />May 8 0.5 5.8 5.2 <br />May 15 8.4 3.6 3.4 <br />May 22 29.0 7.6 6.2 <br />May 29 30.9 6.8 6.8 <br />Jun 5 53.7 7.6 14.9 <br />Jun J.2 124.1 19.2 26.2 <br />Jul 10 95.8 <br />~/ Data from Nance (1997) ; mean values were rounded. <br /> ########## <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />the literature of benthos in running waters (Hynes 1970; Welcomme 1985) <br />that document a higher species diversity and often larger numbers of <br />benthic macroinvertebrates occur in riffles with large rock, cobble, or <br />gravel substrates than river reaches with sand and silt substrates. <br />Benthic invertebrates that inhabit riffles colonize downstream reaches by <br />movement through the substrate, displacement by high streamflows, and <br />drift. However, upstream movement is hindered by long reaches of sand- <br />silt substrates and streamflows 12 cm/s or greater (Luedtke and Brusven <br />1970). Generally, upstream movement by benthic invertebrates is 5 to 30% <br />of the downstream drift (Bishop and Hynes 1969). Benthic invertebrates <br />produced in floodplain habitats move or are carried by currents and can <br />provide a substantial part of the food base for fish in the main channel <br />and backwaters (Eckblad et al. 1984). <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />Although species diversity of benthic invertebrates in low velocity <br />habitats with sand and silt substrates of the Upper Colorado River Basin <br />is less, the numbers of certain taxa (e.g, chironomids) can be very high. <br />Chironomids constitute a significant part of the diets of larger larvae, <br />juveniles, and even adult endangered fishes in low velocity habitats of <br />the Upper Basin so that taxon is emphasized in this discussion of benthic <br />production. The upper part of the range for chironomids was 4,150 per m2 <br />in the main channel, 31,125 per m2 in backwaters, and 23,055 per m2 in <br />floodplain habitats of Upper Basin rivers (Table 4). More detail of the <br />mean numbers of chironomids in different habitats of the Upper Basin are <br />summarized in Appendices 1 and 2. <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />The numbers of chironomids, found in the main channel of the middle Green <br />River, ranged between 360 to 4,645 organisms/m2 and the numbers ranged <br />from 4,820 to 28,860 organisms/m2 for backwater habitats in 1987 (Appendix <br />1; Grabowski and Hiebert 1989). In 1988, Grabowski and Hiebert reported <br />the range in numbers of chironomids between 280 and 5,000 organisms/m2 for <br />the main channel of the river and between 3,330 and 28,890 organisms/m2 in <br />backwater habitats. Obviously, the backwater habitats were more <br /> <br />I <br /> <br />I <br /> <br />14 <br /> <br />I <br />I <br />