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<br />I <br /> <br />I <br /> <br />1987 and 1988. Mabey and Shiozawa (1993) documented zooplankton densities <br />in the middle Green River as 0.3 to 1.3 organisms per liter, 1.5 to 7.1 in <br />the Ouray backwater, 63.4 at Intersection Wash (a large backwater), and <br />206 to 690 in an floodplain depression (Old Charley Wash) along the middle <br />Green River (Appendix 1) . <br /> <br />I <br /> <br />I <br /> <br />In an open water habitat of the Moab Slough along the Colorado River <br />immediately upstream from Moab, Utah, the density of cladocerans and <br />copepods averaged about 2 per liter for backwater sites and about 36 per <br />liter for an open water wetland during the summer of 1993 (Appendix 2; <br />Cooper and Severn 1994a). Cooper and Severn did not collect any <br />zooplankton during their sampling from the main channel in this reach of <br />the Colorado River. Samples of cladocerans and copepods from the <br />Escalante Ranch site on the middle Green River, upstream from Jensen, <br />Utah, contained 0 per liter for the main river channel, a mean of 41 per <br />liter for backwaters, and a mean of 71 per liter for an open water wetland <br />(Appendix 2; Cooper and Severn 1994b). The mean number of cladocerans and <br />copepods from a backwater of the Gunnison River at the Escalante State <br />Wildlife Area, about 5 miles downstream from Delta, Colorado, was 11 per <br />liter and the mean number from an open water wetland was 25 per liter <br />(Appendix 2; Cooper and Severn 1994c). Cladocerans and copepods samples <br />from a floodplain depression (Old Charley Wash) in the middle Green River <br />on the Ouray National Wildlife Refuge contained a mean of 31 per liter in <br />1993 (Appendix 2; Cooper and Severn 1994d). Samples taken from the main <br />channel and a backwater on the refuge did not contain any cladocerans or <br />copepods when sampled (Appendix 2; Cooper and Severn 1994d). The density <br />of cladocerans was 26 per liter and the density of copepods was 28 per <br />liter in a gravel-pit pond along the Colorado River in October, 1993 <br />(Appendix 2; Cooper and Severn 1994e). These studies document that the <br />highest zooplankton densities in the Upper Basin were in floodplain <br />habitats. Seasonal dynamics of zooplankton were not made in the studies <br />summarized above. <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />The mean number of aquatic organisms captured in 5 vertical tows with a <br />20.3 cm plankton net in backwaters of the lower Green River in 1997 varied <br />between 2.4 to 124.1 organisms (combined zooplankton and free-swimming <br />benthic invertebrates) per liter, depending upon site and date of <br />collection (Table 3; Nance 1997). The mean numbers of aquatic organisms <br />increased in the lower Green River between May and July that corresponded <br />to warmer water temperatures (Table 3). The four major taxa represented <br />in lower Green River samples were Nemotoda, Oligochaeta, Rotifera, and <br />Copepoda. Several of the minor taxa included Cladocera and Chironomidae. <br /> <br />I <br /> <br />I <br /> <br />Microcrustaceans that form zooplankton communities are also found in <br />substrates of aquatic habitats as benthos. Mabey and Shiozawa (1993) <br />reported between 1,000 and 6,300 benthic microcrustaceans (Cladocera, <br />Copepoda, and Cyclopoida) per square meter in the main channel of the <br />middle Green River through the Ouray National Wildlife Refuge, between <br />4,900 and 6,000 organisms per square meter for a backwater, 23,000 per <br />square meter at Intersection Wash and between 8,600 and 263,000 per square <br />meter in Old Charley Wash. The mean number of aquatic organisms collected <br />in five 1.85 cm bottom cores from backwaters in the lower Green River <br />varied from 65,200 to 562,200 organisms per square meter, depending upon <br />the site and date of collection (Nance 1997). <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />C. Production of Benthic Macroinvertebrates in Various Aquatic Habitats. <br />Various factors affect the microdistribution and habitats of benthic <br />invertebrates in streams including water velocity and substrate size such <br />as rock, cobble, gravel, silt, and detritus (Rabeni and Minshall 1977). <br />The experimental results of Rabeni and Minshall (1977) are supported by <br /> <br />I <br /> <br />I <br /> <br />13 <br /> <br />I <br />