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<br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I' <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br /> <br />I <br />I <br /> <br />Table 4. Range of the mean number of chironomids from investigations of <br />various habitats in the Upper Colorado River Basin. 1/ l/ <br /> <br />Habitat Range of the mean number of organisms/m2 <br /> <br />Main Channel <br /> <br />20 <br /> <br />4,150 <br />31,125 <br /> <br />Backwaters <br /> <br />83 <br /> <br />Floodplain Habitats <br /> <br />o <br /> <br />23,055 <br /> <br />1/ The mean number of chironomids were obtained from the following studies: <br />Grabowski and Hiebert (1989); Cooper and Severn (1994a), b, c, d, and e; <br />Wolz and Schiozawa (1995). The main channel, backwaters, and floodplain <br />habitats contained primarily sand and silt at the sites sampled so <br />chironomids were used as a measure of relative abundance of benthic <br />macroinvertebrates. The number of samples taken, sampling gear used, <br />seasons (late spring and summer) and years of sampling differed so ranges <br />are used to illustrate the relative productivity of chironomids. <br /> <br />l/ Most (4) of the investigations were made on the Green River. However the <br />data include two investigations on the Colorado River and one <br />investigation on the Gunnison River, a major tributary of the Colorado. <br /> <br />########## <br /> <br />productive in chironomid production than the main channel. Another study <br />evaluated macroinvertebrate densities in the main channel, side <br />(ephemeral) channel, and two types of backwater habitats in the middle <br />Green River, downstream of Vernal, Utah. The numbers of chironomids in <br />the main channel ranged between 3,500 and 4,200 organismsfm2, the range <br />for the side (ephemeral) channel was between 2,300 and 8,100, in one large <br />backwater (basically a riverside lagoon) the range of chironomids was <br />between 9,000 and 23,000, and in the other backwater, the range was <br />between 22,800 and 31,100 organisms/m2 (Appendix 2; Wolz and Shiozawa <br />1995). Four baseline studies were conducted of wetland habitat sites in <br />bottomlands of rivers in the Upper Basin. The wetland at Escalante Ranch <br />along the middle Green River, upstream from Jensen, Utah, produced a mean <br />of 17 chironomids/m2 in the main channel, 17 organisms/m2 in a backwater, <br />and 31 organisms/m2 for an open water wetland (Appendix 2; Cooper and <br />Severn 1994b). At another wetland site, Cooper and Severn (1994a) <br />reported a mean of 11 chironomids/m2 in the main channel of the Colorado <br />River immediately upstream from Moab, Utah, 4 in a backwater site and 11 <br />in an open water wetland. The Gunnison River at the Escalante State <br />Wildlife Area, about five miles downstream from Delta, Colorado, contained <br />a mean of about 496 chironomids/m2, a backwater contained 1,141 and an <br />open water wetland contained 1,092 (Appendix 2; Cooper and Severn 1994c). <br />The mean number of benthic chironomids/m2 in a floodplain depression (Old <br />Charley Wash) was 33, 21 in a backwater, and 10 from the channel of the <br />middle Green River adjacent to the floodplain on the Ouray National <br />Wildlife Refuge, Utah (Cooper and Severn 1994d) . <br /> <br />The mean number of aquatic organisms (combined species) collected in five <br />1.85 cm bottom cores from backwaters in the lower Green River varied <br />between 65,200 and 562,200 organisms/m2, depending upon the site and date <br />of collection (Nance 1997). <br /> <br />15 <br />