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great for germination. Therefore, it seems logical that multiple -lift <br />topsoiling may increase establishment of volunteer plant species from <br />stored propagules in the topsoil; if such species are desirable from a <br />synecological standpoint, ultimate plant community diversity may conse- <br />quently also be increased. <br />It should be recognized that the above relationship of multiple - <br />lift topsoiling to vegetation diversity is presently at best only a <br />logical hypothesis unsubstantiated by data. However, if the hypothesis <br />is ultimately proved by future research, multiple -lift topsoiling may <br />become a valuable strategy to increase vegetation diversity on mined <br />lands. <br />Topsoil Depth <br />As noted previously, considerable research has been conducted in <br />the Northern Great Plains Region on effects of topsoil replacement depth <br />on total vegetation productivity and /or cover. However, published infor- <br />mation on the influence of topsoil depth on plant community composition <br />and diversity is surprisingly limited and often somewhat contradictory. <br />Pinchak (31) noted no significant differential effects of topsoil <br />depths from 20 to 60 cm on vegetation diversity on uranium mined lands <br />in Wyoming revegetated primarily to perennial grasses. However, a <br />number of other researchers have demonstrated definite differences in <br />establishment /growth among different plant species or growth forms in <br />response to topsoil depth variation. Redente et al. (35, 36), for <br />example, noted shrub productivity to generally and consistently increase <br />with increasing topsoil depth while perennial grass productivity in- <br />creased only to a certain point; forb productivity was actually highest <br />under the shallowest topsoil depth evaluated. Biondini and Redente (4) <br />further noted differences in performance among types of perennial grass <br />species, with bunchgrasses more responsive to increased topsoil depth <br />than rhizomatous grasses. The above authors and Allerdings and <br />Redente (1) actually found highest vegetation diversity under shallowest <br />topsoil depths evaluated, although total productivity was reduced. <br />The above results indicate that topsoil depth can indeed influence <br />plant community composition and diversity, and that diversity and <br />productivity may not always be similarly affected by increasing topsoil <br />depth. They suggest that if adequate diversity within plant communities <br />is an important reclamation goal, topsoil should sometimes be reapplied <br />at depths shallower than those necessary to maximize total productivity. <br />In short, "intermediate" topsoil depths may provide, in many cases, an <br />optimum compromise between diversity and productivity. <br />Topsoil Treatment /Management <br />Although topsoiling may usually be expected to improve initial <br />and /or potential minesoil fertility, numerous studies in the Northern <br />Great Plains have indicated generally positive effects of N and P ferti- <br />lization on vegetation productivity. Consequently, fertilization has <br />become a common reclamation practice in the region. However, while <br />263 <br />