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Site <br />Ungrazed <br />Total <br />Grazed <br />Total <br />Fresh stockpile <br />1- year -old <br />stockpile <br />652 <br />Depth <br />0 -15 3985+ <br />0 -2.5 <br />0 -7.5 <br />'2.5 -7.5 <br />7.5 -15 <br />4660 <br />7240 <br />2580 <br />510 <br />0 -15 7750 <br />0-7.5 <br />Germinable seeds j <br />(cm) Mean 95% confidence limit <br />0 -2.5 2235+ 1465 -3000 58 2.99 Artemsia dracunculus <br />0 -7.5 2980+ 1960 -4005 3.02 Melilotus officinalis <br />'2.5 -7.5 745+ 16 Bouteloua gracilis <br />7.5 -15 1005 60 -1950 26 1.79 = Conyza canadensis <br />- y'. =' Hedeoma hispida <br />3195 -6120 <br />4990 -9490 <br />0 -1065 <br />255++ 48362 <br />0 -7.5 520 ++ 113 -927 <br />'calculated by difference of 0-2.5 and 0.75 cm levels, confidence limits could not be calculated <br />+significantly fewer seeds than equivalent depth level on grazed site P5.05). - <br />+ +number of seeds on stockpiles not significantly different from each other but significan <br />Literature Cited <br />Barton, L.V. 1962. The germination of weed seeds. Weed Sci. 10:174 -182. <br />Barzaz, F.A. 1970. Secondary dormancy in the seeds of the common ragweed Ambro- <br />sia artemisafolia. Bull: Torrey Bot. Club 97:302 -305. <br />Bazzaz, F.A. 1979. The physiological ecology of plant succession. Ann. Rev. Ecol. <br />Syst. 10:351 -371. ' <br />Beauchamp, H., R. Lang, and M. May. 1975. Topsoil as a seed source for reseeding <br />strip mine spoils. Res. J. 90, Agr. Exp. Sta., Univ. of Wyoming. - ` = <br />Best, K.F., J. Looman, and J.B. Campbell. 1971. Prairie grasses. Can. Dept. Agr. <br />Publ. 1413. 239 p. <br />Brenchley, W.E. 1944. Weed control and the dormancy of weed seeds. J. Ministry of <br />Agr., United Kingdom 50:452 -455. <br />Brenchley, W.E., and K. Warington. 1930. The weed seed population of arable soiL 1. <br />Numerical estimation of viable seeds and observations on their natural dormancy. <br />J. Ecol. 18:235 -272. <br />Brenchley, W.E., and K. Warington.1933. 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Longevity, periodicity of germina- <br />tion, and vitality of seeds in cultivated soil. J. Agr. Sci. 26:307 -349. <br />Chippindale, H.G., and W.E.J. Milton. 1934. On the viable seeds present in the soil <br />beneath pastures. J. Ecol. 22:508 -531. <br />Connel, J.H., and R.O. Slatyer. 1977. Mechanisms of succession in natural communi- <br />ties and their role to community stability and organization. Amer. Nat. 111. 1119- <br />11 <br />Darlington, H.T. 1931. Dr. W.J. Beals seed viability experiment. Amer. J. Bot. <br />18262 -265. <br />Darlington, H.T. 1941. The sixty -year period of Dr. Beal's seed viability experiment. <br />Amer. J. Bot. 28:271 -273. <br />Dunham, R.S., A.H. Larson, and R.G. Robinson. 1947. Weed seedlings. Minn. Agr. <br />Exp. Sta. Bull. 397:1 -36. <br />Dwyer, D.D., and E. Aguirre V. 1978. Plants emerging from soils under three range <br />condition classes of desert grassland. J. Range Manage. 31:209 -212. <br />Gleason, H.A., and A. Cronquist. 1963. Manual of vascular plants of northeastern <br />United States and adjacent Canada. Van Nostrand Reinhold Co., New York. 810 p. <br />59 <br />35 <br />6 <br />Seed Important species <br />diversity (>5% of total) <br />3.63 <br />,.3.05 <br />1.25 <br />Conyza canadensis <br />Festuca octollora <br />Hedeoma hispida <br />Cerastium brachypodum <br />Plantago patagonica <br />2.32 Andropogon gerardi <br />A. scoparius <br />Agrostis scabra <br />Bouteloua gracilis <br />Festuca octollora <br />Artemesia frigida <br />Conyza canadensis <br />Androsace occidentalis <br />Bouteloua gracilis <br />. Festuca octoflora <br />% of <br />total <br />28 <br />16 <br />8 <br />5 <br />5 <br />19 <br />14 <br />13 <br />10 <br />8 <br />20 <br />20 <br />20 <br />20 <br />20 <br />22 <br />22 <br />11 : -- <br />11 <br />11 <br />nngrazCd'sites (0 cm; P5..00!) <br />es ais vegetation processes. John Wiley, Chichester, <br />role ofpioneering specks on the reclamation of North Dakota` <br />Dissertation; Univ. of North Dakota, Grand Forks, <br />all.`1982. Rec mation of coal - mined lands: the role,of <br />attd other pioineert; Rec. and Re veg. Res. I:123 k <br />196ECootent efburied seeds in arable soil in Denmark and its relation <br />ulatiorr. Dansk Botariisk Arkiv 27:1 • i ' _ <br />Johoat¢o; -`X.; a' wulik :; ?W . Stringer, 1968. Viable seed po pulations in <br />Alberti peaide fopaotls. ,Cu 1 P1 Sci. 49 :75 - 82. - - <br />Uppert. R.D, and H.H. Hoptloa.1950. Study of viable seeds in various habitats in <br />nixed prsitrie: Kan'Aead. Sci,' Trans. 53:355 -364. -_ < % - <br />Major, Jy aid W.T.I'yott.1966..Buried, viable seeds in two California bunchgrass <br />sites and their bearing on the definition of a flora. Vegetatio 13:253 - 282.', . -= <br />�►1eRil1, M.,and G.R: Segar 1973; Earthworms and seeds. Nature 24 -482, , - <br />Milton W.E.L 194*. Theburied viable seed content of upland soils in Montgomery' - <br />• =' shire. Empire t Exp Agr `16163- 177. Mortimer, 10 Studies of germination and establishment of selected species <br />- with special reference to the of Ph.D. thesis, University College of North <br />'f=' Wales, 'Bangor; '?„h tt Z ` _ -• `t.. - <br />Noble, I.R , and R.0 Shafer. 1988. The use of vital attributes to predict successional <br />changes in plant,communities subject to recurrent disturbances. Vegetatio 43:5 -21. -- <br />Prince, F.S., and A.1R. Hodgdon: 1946. Viable seeds in old pasture soils. N.H. Agr. - <br />Exp. "Sts. Tech.'BuII. 89:3 16. ! `: ' ' - . <br />Rabinowitz, D. 1981. Buried viable seeds in a North American tall -grass prairie: the <br />resemblance of their abundance and composition to dispersing seeds. Oikos 36:191- <br />Roberts, H.A. 1963. Studies on the weeds of vegetable crops. 111. Effect of different <br />primary cultivations on the weed seeds in the soil. J. Ecol. 51:83 -95. " ' <br />Roberts, H.A. 1968. The changing population of viable weed seeds in an arable soil. <br />. - - Weed Res. 8:253 -256. <br />Roberts, H.A., andJ.E.Nellson.1981. Changes in the soil seed bank of four long -term <br />crop /herbicide 1 appl. Ecol. 18:661 -668. <br />Robinson, R.G. 1949. Annual weeds, their viable seed population in the soil, and their <br />effect on yields of oats, wheats, and flax. Agron. J. 41:513 -518. _ . <br />Stevens, O.A. 1963. North Dakota plants. ND Inst. for Regional Studies, Fargo, ND. <br />324 p. <br />Thompson, K., and 32. • Grin e.'1979. Seasonal variation in the seed banks of <br />herbaceous ° species in ten contrasting habitats. J. Ecol. 67:893 -922. <br />Thurston, J.M. 1960. Dormancy in weed seeds. Pages 69-82 in Harper, J.L. ed. The <br />Biology of Weeds. Blackwell Sci. Pub., London, U.K. <br />Van Bruggen, T.1976. The vascular plants of South Dakota. Iowa State Univ. Press, <br />Ames. 538 p. " - <br />Whitman, W.C. and M.K. Wag, 1975. 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