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and water quality after wildfires (e.g., Tiedeman et al., <br />1978; Robichaud et al., 2000), but much less is known <br />about the effects of fires on channel morphology and <br />aquatic habitats. The effects of fires on water quality <br />are highly variable, as the type and magnitude of change <br />depends on the size and severity of the fire, site char- <br />acteristics such as slope and soil type, vegetation type, <br />amount and type of precipitation, and the rate and type <br />of regrowth (Friedrich, 1951). Because there are so many <br />different processes operating at different time scales, it <br />is difficult to compare results from different watersheds <br />(Krammes, 1960). The seemingly disparate results can <br />only be resolved by understanding how each fire affects <br />the underlying processes of runoff, erosion, and biogeo- <br />chemical cycling. <br />By consuming organic matter, both controlled and un- <br />controlled fires cause some nutrient loss. These losses <br />are due to nutrient volatilization, particularly in the case <br />of nitrogen, and nutrient leaching. On the other hand, <br />most fires alter the soil environment in ways that lead to <br />increased microbial activity. The conversion of organic <br />matter to ash and the increase in microbial activity usual- <br />ly results in a short-term increase in nitrogen availability <br />and movement. Depending on the monitoring frequency <br />and site conditions, an ammonium pulse may occur <br />before the more commonly observed nitrate pulse. The <br />short-lived increase in nitrogen availability may help <br />stimulate any surviving vegetation or the establishment <br />and regrowth of new vegetation. <br />Soil erosion rates may increase after fires, but the magni- <br />tude and persistence of an increase is highly variable and <br />dependent in large part on fire severity (Tiedemann et al., <br />1979; Robichaud and Waldrop, 1994; Benavides -Solo- <br />rio, 2003). Controlled or prescribed fires typically are <br />designed to remove fuels or selected vegetation classes, <br />or modify a vegetation type. Controlled burns usually do <br />not consume all the protective duff or litter layer over <br />large areas, while high severity wildfires can expose <br />the mineral soil over relatively large areas. Post -fire in- <br />creases in erosion can result from changes in a number <br />of processes, and this can make it difficult to generalize <br />or predict the effects of a given fire. Higher erosion rates <br />after fires can result from: (1) the decrease in litter and <br />vegetative cover; (2) changes in soil properties, including <br />the loss of organic matter, formation of a water repellent <br />layer at or below the soil surface, and sealing of the soil <br />surface; (3) increased rill erosion due to the increase in <br />overland flow; (4) channel incision due to the increase in <br />runoff and decrease in surface roughness; and (5) mass <br />movements. Soil erosion can reduce the amount of soil <br />nutrients on site, but more nutrients are usually lost by <br />28 <br />burning the vegetation unless post -fire soil erosion rates <br />are very high. <br />Some studies in the Colorado Front Range have found <br />very little erosion after wildfires (Delp, 1968; Meyers, <br />1968), while erosion rates after the 1996 Buffalo Creek <br />fire increased by several orders of magnitude (Moody <br />and Martin, 2001). Morris and Moses (1987) measured <br />large increases in surface erosion after several fires in the <br />Colorado Front Range, while Cannon et al. (1997) docu- <br />mented an increase in mass movements after the Storm <br />King fire near Grand Junction. A series of detailed stud- <br />ies on the June 2000 Bobcat fire just west of Fort Collins <br />are showing that sites burned at high severity produce <br />much more sediment than sites burned at moderate or <br />low severity. Rainfall simulations on 1 m2 plots indicate <br />that sites burned at high severity produce 10 -30 times <br />as much sediment as unburned sites, and sites burned at <br />moderate severity produced 2 -6 times as much sediment <br />as unburned and low severity sites (Benavides - Solorio <br />and MacDonald, 2001; Benavides - Solorio, 2003). Moni- <br />toring of sediment production at the hillslope scale from <br />the Bobcat and several other fires confirm the large dif- <br />ferences in sediment production with fire severity (Bena- <br />vides- Solorio, 2003). <br />Data from the Buffalo Creek, Bobcat, and other fires <br />indicate that at least 80% of the annual erosion is driven <br />by summer convective storms with rainfall intensities of <br />10 mm (0.4 inches) per hour (Moody and Martin, 2001; <br />Benavides - Solorio, 2003; Kunze, 2003). In the case of <br />the Buffalo Creek and Bobcat fires there were unusu- <br />ally large convective rainfall events in the first one or <br />two summers after burning, but similar storms did not <br />occur after the Hayman fire. It has been suggested that <br />large burned areas may increase the likelihood of large <br />convective storms, and this could be an important topic <br />for further study. <br />Another important issue is the rate at which sites recover <br />following burning. Plot -scale studies on the Buffalo <br />Creek fire indicated that erosion rates on burned hill - <br />slopes were not significantly different from unburned <br />hillslopes by the fourth year after burning. Data from <br />rainfall simulations and sediment fences confirm that <br />runoff and erosion rates six years after the 1994 Hour- <br />glass fire were not substantially different from the rates <br />observed from unburned sites and areas that had been <br />burned at low severity (Benavides - Solorio, 2003). How- <br />ever, at the larger scale channel incision and the deposi- <br />tion of sediment in downstream areas may be much more <br />persistent. Hence the recovery of hillslope -scale pro- <br />cesses may be relatively rapid due to vegetative regrowth <br />