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downstream? (3) What change in flow is detectable? and <br />(4) Can changes in forest cover cause changes in flows <br />on larger basins, even if the change in flow is not statis- <br />tically detectable? The answers to these questions have <br />important implications with respect to the larger -scale <br />changes in runoff that are of most interest to resource <br />managers and the public. <br />The answer to the first question — how much of a basin <br />must be treated in order to detect a change in runoff — is <br />relatively well known. A 1982 review of paired catch- <br />ment experiments concluded that at least 15 -20% of a <br />forested basin must be treated within a short time period <br />in order to detect a change in runoff (Bosch and Hewlett, <br />1982). Troendle and Leaf (1980) noted that 20 -30% of a <br />watershed must be treated to detect a statistically signifi- <br />cant change in flow, and this value has been supported <br />by the magnitude of flow changes observed in paired <br />watershed experiments in the Rocky Mountains (Table <br />2.1). At Coon Creek in south - central Wyoming there was <br />a detectable change in annual water yield as a result of <br />harvest and road - building on 24% of the watershed (Tro- <br />endle et al., 2001). Small increases in annual water yields <br />were observed by removing 31 -33% of the trees in two <br />small watersheds in northern Arizona (Baker, 1986). <br />Harvesting 36% of the North Fork of Deadhorse Creek <br />caused a significant change in runoff at the sub - watershed <br />scale, but a change in water yields was not detectable at <br />the main weir because the treatment only removed about <br />5% of the forest canopy above this weir. For ponderosa <br />pine forests in the Black Hills of South Dakota, 25 -30% <br />of the forest canopy may have to be removed in order to <br />obtain a detectable increase in annual water yields. This <br />higher threshold is probably due to the drier conditions <br />and relative absence of a strong snowmelt peak. <br />The second question is whether a flow increase at an <br />upstream location will be delivered to a downstream res- <br />ervoir, diversion intake, or other location of interest. The <br />general principle is that streams are relatively efficient <br />in conveying water downstream, and this is particularly <br />true in headwater forested areas (e.g., Troendle, 1982). <br />However, as streams flow from the mountains and onto <br />the plains streams may become losing rather than gain- <br />ing. This means that groundwater is no longer flowing to <br />the streams ( "gaining "), but streamflow is seeping out of <br />the channel to support the local groundwater table ( "los- <br />ing"). Hence some of an increase in streamflow due to <br />forest management may be transferred to the adjacent <br />groundwater, particularly in larger basins. The availabil- <br />ity of the resultant increase in groundwater for human <br />use will vary from basin to basin. Riparian vegetation <br />19 <br />and surface evaporation also can reduce the amount of <br />streamflow that is delivered downstream. <br />The proportion of a change in streamflow that is trans- <br />mitted to a specific location can vary seasonally and <br />from year to year. During spring snowmelt and early <br />summer streams are more likely to be gaining, while <br />later in the summer groundwater levels tend to drop <br />and streams may shift from gaining to losing (e.g., Graf, <br />1997). Losses to riparian vegetation and surface evapo- <br />ration generally will be much less in spring than later in <br />the summer. Thus an increase in spring snowmelt gener- <br />ally should be more readily translated downstream than <br />an increase in low flows. Some of the increases in peak <br />flows may be lost to bank storage and overbank flows, <br />and this will attenuate high flows in the downstream di- <br />rection. The extent to which an increase in flow is trans- <br />lated downstream will depend on the conditions within <br />that basin, but generally one would expect greater losses <br />with increasing spatial scale and increasing distance <br />from the mountain front. <br />The third issue is the detectability of a given change. <br />The ability to detect change will vary according to the <br />accuracy of the discharge measurements, the magnitude <br />of the imposed change, the variability of the data, the <br />length of the record prior to and after treatment, the cer- <br />tainty with which we want to detect change (i.e., level of <br />significance), and the certainty of detecting change when <br />in fact there is a change (i.e., power) ( Loftis et al., 2001). <br />For small research catchments discharge is typically <br />measured at carefully- designed weirs with impermeable <br />cutoff walls, and in these situations the uncertainty in <br />discharge is generally believed to be on the order of only <br />2 -3 %. In contrast, the uncertainty of a single flow mea- <br />surement in a natural channel is generally believed to <br />be around 5% for moderate flows (Kennedy, 1983) and <br />substantially larger at higher flows (Dickinson, 1967). <br />The extrapolation from individual flow measurements <br />to seasonal or annual water yields may involve a further <br />degradation in accuracy because of the greater error in <br />measuring the highest flows, the uncertainty in the re- <br />lationship between stage and discharge, and the greater <br />potential for missing data (Kennedy, 1983). Hence the <br />uncertainty of flow measurements at a typical stream <br />gauging station is probably closer to 10 %, or several <br />times greater than in a research setting, and this greater <br />uncertainty limits our ability to detect a change in runoff <br />due to a given change in vegetation or land use. <br />The use of paired- watershed design greatly increases <br />our ability to detect change (Wicht, 1967; Loftis et al., <br />2001), as the data from a control watershed can be used <br />