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become partners in the project, including the Northern Colorado Water Conservancy District (NCWCD), Denver Water, <br />and the U.S. Bureau of Reclamation, Aspinall Unit (Gunnison Basin). <br />The physical basis for the reconstructions <br />Tree growth is primarily controlled by climate conditions during the year prior to and including the growing season. At <br />lower elevations in Colorado, variations in tree growth closely reflect the amount of soil moisture at the onset of the grow- <br />ing season, which is controlled by variations in precipitation, and, to a lesser degree, temperature, humidity, and evapora- <br />tion. Since annual streamflow integrates both precipitation and temperature over the course of the previous seasons (simi- <br />lar to tree rings), it is closely correlated with annual tree growth. Trees that provide the best information about streamflow <br />variability —those particularly sensitive to variations in moisture — include species such as ponderosa pine, pinyon pine, <br />and Douglas -fir, growing in open stands on dry and rocky sites where soil moisture storage is minimal (Figure 2). Trees <br />growing in these sites are also less likely to be subject to non - climatic disturbances, such as fires and insect infestation, and <br />the effects of competition from nearby trees. In addition, the oldest individuals (up to 900 years old) of these species tend <br />to be found on these sites. <br />Developing the reconstructions <br />Tree -ring reconstructions of streamflow are developed from multiple <br />tree -ring chronologies. A tree -ring chronology is a time - series of annual <br />values derived from the ring -width measurements of 10 or more trees of <br />the same species at a single site. To create a tree -ring chronology, cores <br />from the sampled trees at each site are crossdated (patterns of narrow <br />and wide rings are matched from tree to tree) to account for missing or <br />false rings, so that every annual ring is absolutely dated to the cor- <br />rect year. Then all rings are measured to the nearest 0.001mm using a <br />computer- assisted measuring device. After age - related trends in growth <br />are statistically removed, the ring -width values from all sampled trees <br />for each year are averaged to create a time series of annual ring -width <br />indices. The complete series of ring -width indices from a site is called a <br />tree -ring chronology. <br />Once a gaged natural flow record of interest is selected for reconstruc- <br />tion, a set of chronologies from the region near the gage is calibrated <br />with the gage record to form a reconstruction model. A statistical <br />technique called multiple linear regression is commonly used. The <br />reconstruction is evaluated by comparing the observed gage values with <br />the reconstructed values and assessing the amount of variance in the <br />gage record that is explained by the reconstruction. The reconstruction <br />model is then validated by testing it on a portion of the gage data that <br />was withheld from the calibration process. <br />What the reconstructions show <br />So far, we have developed <br />reconstructions, extending back <br />to the 1400s to 1600s, of the 15 <br />stream gages that comprise the <br />water supply systems for Denver <br />Water, NCWCD, and the city of <br />Boulder (Figure 1). Our tree - <br />ring reconstructions match the <br />variability in the gaged records <br />of streamflow very well (Figure <br />3). Statistically speaking, 60% <br />to 75 % of the variance in the <br />gaged records is explained by the <br />0 400000 <br />w <br />d <br />u 300000 <br />0 <br />E 200000 <br />E <br />L <br />V) 100000 <br />3 <br />C <br />0 <br />a <br />Figure 2. Sampling a 400 - year -old ponderosa pine <br />near Deckers, Colorado <br />1910 1920 1930 1940 1950 1960 1970 1980 1990 2000 <br />Figure 3. Blue River at Dillon Reservoir undepleted gaged annual flow,1916 -2002 (light) and <br />tree -ring reconstructed flow (dark). Note the close fit of the two time series (M = 0.63), especially <br />for the most extreme low -flow events (1954, 2002). <br />29 <br />