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<br />1724 <br /> <br />BESTGEN ET AL. <br /> <br />Biotic Controlling Factors <br /> <br />Embryos and Larvae <br />in substrate <br /> <br />Abiotic Controlling Factors <br /> <br />Number and condition of adu~s }- -{Timing and magnitude of runoff <br />Age and size strueture of adu~s Substrate characteristics, siltation, oxygen <br />Timing of spawning . Discharge fluctuations <br />Predation on embryos and larvae Water temperature <br />Physical disturbance of spawning site by other fish Stochastic events (e.g., floods and pollutants) <br /> <br />Dispersing Larvae <br />in main channel <br />and channel margins <br /> <br />Predation <br />Starvation <br />Habitat selection <br /> <br />Magnitude and duration of spring runoff <br />Water temperature and variability <br />Geomorphic factors <br />Turbidity <br /> <br /> <br />Predation, affected by individual characteristics r ~ <br />Growth rate Discharge- and geomorphic-related habitat availability <br />Condition and starvation Discharge fluctuations and habitat stability <br />Food abundance Water temperature <br />Intra- and interspecific competition Stochastic events (e,g,. floods and pollutants) <br /> <br /> <br />Size-related energy stores <br />Size-related swimming ability <br />Food abundance <br />Habitat-related movement patterns <br /> <br />-[Discharge-related habitat availability <br />Discharge fluctuations and habitat stability <br />Water temperature <br /> <br />Growth <br /> <br /> <br />Predation, affected. by individual characteristics }- -[spring floodplain connectivity <br />Food abundance Main channel discharge-dependent seasonal habitat <br />Intra- and interspecific competition availability <br />Habitat-related movement patterns Water temperature regimes <br />Reproductive condition Stochastic events (e.g., pollutants) <br />Drought effects on condition and survival <br /> <br />FiGURE I.-Conceptual life history model of Colorado pikeminnow recruitment to various developmental stages and important <br />biotic and abiotic factors that control them. <br />