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31 <br />The filling period was characterized by lower peak flows while • <br />temperature remained essentially unchanged, thus reproductive failure., <br />was attributed to flow regulation. <br />Resource Consultants, Inc. (1985) pointed out that bonytail chub <br />populations declined earlier in the Upper Colorado River mainstem than <br />in the Green River, and that large scale flow depletions occurred earlier <br />in the Colorado River mainstem than in the Green River. <br />Razorback sucker <br />o Recruitment of razorback sucker is highest in high flow years. <br />Johnson and Rinne (1982) reported a correlation between high flow levels and <br />reproductive success of razorback sucker in the lower Colorado River Basin. • <br />It should be noted, however, that razorback sucker populations in the lower <br />basin are often associated with lacustrine habitats, thus extrapolation of such <br />data should be done with caution. <br />Seasonal flow needs. <br />Colorado squawfish <br />o Spring peak flows approaching natural (historic) levels are best for Colorado <br />squawflsh spawning for all but very high discharge years. <br />Haynes et al. (1985) reported the ideal spring flow regime to be one of <br />relatively high peak discharge, but Jones and Tyus (1986) and Tyus et •