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<br />" <br /> <br />consequence of the Yampa's ameliorating effect on water temperature. <br />Work by the Utah Cooperative Fishery Research Unit during 1963-1966 <br />focused on basic life history of the Colorado squawfisn and bony tail {Vanicek <br />1967; Vanicek and Kramer 1969; Vanicek et ll.--1970Yand on macroinvertebrate <br />abundance and distribution {Pearson 1967; Pearson et ll. 1968}. These <br />studies suggested that year classes of Colorado squawfish were strong in <br />1959~ 1961, 1963, 1964, and possibly 1966 (Vanicek 1967), but weak in 1962 <br />and 1965. Formation of strong year classes during years when streamflow and <br />water temperatures are more suitable for survival of_ recruits could be an <br />evolutionary life history strategy of this species (Tyus 1986, Chapter 19). <br />Vanicek (1967) also reported difficulty in separating the various species of <br />Gila, especi ally duri ng early 1 ife stages. Thi sled to a study of Gil a <br />taxonomy by Holden (1968; Holden and Stalnaker 1970), but taxonomic ~uestions <br />were not fully answered. It was not until 1989 that the USFWS contracted <br />wit~ the Smithsonian Institution to comprehensively review the problem with <br />the goal of resolving the taxonomic confusion surrounding the Gila species. <br />Holden (1973) studied relative abundance and distribution of native <br />fishes in the upper basin and documented problems with recruitment of <br />squawfish in Echo Park (DNM) where juveniles were abundant in 1968, scarce in <br />1969, and non-existent in 1970. However, he reported young-of-year from the <br />Green River in Desolation Canyon in 1971 and in Canyonlands National Park in <br />1970 and 1971. During 1974-76, studies focused on obtaining further life- <br />history information on the endangered fishes (McAda 1977; Seethaler 1978). <br />McAda (1977; McAda and Wydoski 1980) provided new biological information and <br />synthesized all known life-history data on razorback sucker in the upper <br />basin. Seethaler (1978) provided a comprehensive summary of the life history <br />of the Colorado squawfish, including the first data on reproduction, <br />