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there was a clear relationship between poor year class development in most. <br />native species studied and the artificial hydrograph. <br />Before impoundment behind Missouri River reservoirs .(1926 to 1952), peak <br />discharges generally-occurred in April, and then-again with a larger peak in <br />June (Pflieger and Grace 1987)._ Today,, dam operations reduce flows from April <br />to July for flood control, and increase flows from July to April for <br />navigation, water supply, and hydropower: In addition to such seasonal .shifts <br />in the flow patterns, main stem dams operating for daily hydropower needs-can <br />cause daily fluctuations in water levels in tailwater areas by as much as 2 to <br />3 m (6.6 to 9.8 ft)`. This fluctuation can disrupt the macroinvertebrate <br />community and°larval fish rearing-areas for many miles downstream of the. dam <br />by alternately flooding°and dewatering habitats. <br />Modde and Schmulbach (1973) observed that factors affecting shovelnose <br />sturgeon prey availability within the unchanneli-zed Missouri River include <br />temperature, seasonal recruitment, and changes in density influenced by the . <br />timing and discharge rates from Gavins Point Dam. They hypothesized that the <br />reduction in numbers of shovelnose sturgeon may be due to reduced availability <br />of prey species caused. by high discharges from Gavins Point Dam. <br />Before .the Missouri River was channelized and impounded, it annually eroded <br />3.1 hectares/km of its floodplain (U.S. Army Corps of Engineers 1981). -Most <br />of this erosion has stopped due to channelization and impoundment. Erosion. <br />was a natural funct-ion of the river system,-and through erosion, inorganic <br />sediments, organic matter,. and large woody debris were introduced into the <br />river. This material import was essential to the habitat dynamics and <br />nutrient cycling of the river system.- Such sediment and nutrient discharge <br />are the raw materials for habitat development in the Missouri and Mississippi. <br />River system. Construction of dams eliminated 80 percent of this material. <br />Fremling (1989) reports that the sediment load of the middle Mississippi River <br />has declined 66 percent from pre-1935 levels, mainly due to sediment- <br />entrapment in Missouri River impoundments. This lack of sediment delivery- <br />upset the natural channel equilibrium and was replaced by a variety of <br />nonequilibrium processes such as hydraulic sorting and bed paving, which <br />eventually will eliminate all sediment movement. This has already occurred to <br />some extent and has resulted in reduced bed roughness-and, therefore, reduced <br />substrate diversity. This has. reduced the reproductive success of substrate <br />spawners, such as sauger (Stizostedion canadense), sturgeon, and paddlefish <br />(Hesse and Mestl 1993b). <br />The turbidity caused by suspended sediment also provided the pallid sturgeon <br />and othew native fish, adapted to living in a nearly sightless world,..with- <br />cover:while moving from one snag or undercut bank to another. Today,-water <br />clarity has increased dramatically,-and this essential cover is gone.. Under <br />such conditions,, predation by sight-feedsng predators, such as northern pike- <br />(Esox-Lucius), walleye (Stizostedion vitreum), and.smallmouth bass <br />(raicropterus dolomieui); can be expected to significantly impact native. <br />species not equipped by errolution with good eyesight. <br />12 <br />