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rom a ), and Colorado squawfish. Together these three species comprise 93 - 95% <br />of all backwater fishes (Haines and Tyus 1990). All three species consume <br />chironomid larvae, and the red shiner and young Colorado squawfish also consume <br />zooplankton, and semi-aquatic insects (Catherine A. Karp, personal communication). <br />METHODS <br />Study area <br />The Green River originates in Wyoming and is part of the Colorado River <br />drainage,joining the Colorado River in southeastern Utah. 404 km above its confluence <br />with the Colorado River it enters Ouray National Wildlife Refuge (ONWR). When it <br />reaches ONWR it has a drainage area of 47,723 km2 with an average high flow of <br />11,926 cfs and an average low of 3,530 cfs (U.S.G.S. data). All backwaters used in <br />this study were located on ONWR and were shallow (approximately 0.75 m to 1.0 m <br />deep) with no measurable current except when the wind blew. Secchi disk visibility <br />averaged 22 cm, and the substrate was composed of soft sediments under a thin <br />(approximately 4 cm) layer of flocculent material. <br />Experimental design <br />A randomized block design was used with three treatments per backwater. The <br />treatments were: (a) a four m2 (2 x 2 m) control area, marked by posts, which was <br />open for foraging to all backwater fishes, (b) a four m2 caged area that excluded all <br />fish, and (c) a four m2 cage with 2.5 cm wide by ten cm high (spaced approximately <br />ten cm apart) perforations cut into all four sides. These perforations excluded large <br />fish such as carp (Cyprinus carpio) and channel catfish (Italurus u a us) but <br />4