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INTRODUCTION <br />The role of biotic interactions in structuring natural communities has been well <br />studied (Connell 1983, Schoener 1983, Sih et at. 1985). Investigations of freshwater <br />lentic habitats, especially pelagic, have revealed that predator induced trophic <br />cascades are often a driving force in the communities (Carpenter et at. 1987, <br />Johannes et at. 1989, McQueen et at. 1989, McQueen 1990, Shapiro and Wright <br />1984). However, studies of freshwater lotic communities have had mixed results. Lotic <br />communities with coarse substrates have failed to show significant invertebrate density <br />effects from vertebrate predation (Allan 1982, Reice 1983, Flecker and Allan 1984, <br />Culp 1986, Reice and Edwards 1986), while invertebrates in pools and soft substrates <br />of small streams have responded to fish predation (Wilzbach et at. 1986, Schlosser <br />and Ebel 1989, Angermeier 1985, Gilliam et at. 1989). However, few investigations <br />have tested predatory effects of fishes on invertebrate communities of large rivers. In <br />this study we examined the role of fish predation on the structure of invertebrate <br />communities of the Green River, a large river in eastern Utah. <br />Backwaters are defined as shallow ephemeral embayments adjacent to the main <br />river channel with no measurable water velocity (Tyus and Haines 1991). At least <br />eight native, and fifteen non-native fish species occupy backwaters of the Green River <br />from mid-summer through fall (Haines and Tyus 1990). One of the native fish species, <br />the Colorado squawfish (Ptychocheilus lucius), uses the backwaters as a nursery area <br />(Haines and Tyus 1990). The most abundant backwater fish species are (in <br />descending order) red shiner (Cyprinella lu n i , fathead minnow (Pimephales <br />3