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<br />effects like delay of spawning due to colder water temperatures. In general, however, <br />water quality effects other than temperature have been studied little. Biotic factors are <br />most likely related to predation or competition from nonnative fishes, but may also <br />~ include food supply. <br />Environmental factors that regulate the abundance of a life history stage, or a <br />population, are considered °limiting factors." There are biotic and abiotic factors that <br />regulate growth and mortality, and the relative importance of these limiting factors may <br />vary in time (e.g., with season or with life history stage) or space (habitat occupied by a <br />~ particular life history stage at a particular time of year). Especially for species that are <br />endangered, and thus rare, it may be difficult to define rigorously the factors limiting <br />population size. A certain amount of inference based on best professional judgment <br />therefore becomes necessary. <br />~ Identifying limiting factors is the first step in developing plans for enhancing recovery of <br />the endangered fishes. The next step is assigning priorities for alleviating the <br />limitations imposed by each of the factors. Ideally, priorities for management actions <br />should be established based on the number of fish that will be added to the population. <br />It probably will not be sufficient to address problems one at a time, because multiple <br />~ factors may be acting in concert. <br />In the review that follows, limiting factors are examined on two overlapping scales: <br />basin-wide and species-specific. Limiting factors that exist on the basin-wide scale will <br />affect some or all life history stages of one or more of the endangered fishes. These <br />~ include abiotic factors such as flow and temperature (which are linked to some extent), <br />and biotic factors such as the abundance of nonnative fishes. Basin-wide factors will <br />be reviewed separately because it is easier to understand their origins and the <br />pervasiveness of their effects. It is also necessary to examine limiting factors from a <br />species-specific perspective because it provides the temporal and spatial focus <br />necessary for developing recovery plans efficiently. For each species, limiting factors <br />~ will be reviewed in the context of life history because limiting factors may be different at <br />each stage, especially where the stages occupy different habitat (e.g., Colorado <br />pikeminnow). <br />Basin wide Limiting Factors <br />Abiotic <br />The construction of dams and diversion structures in the Colorado River basin has <br />converted much riverine habitat into reservoirs and smaller lacustrine habitat. Loss or <br />~ alteration of habitat has been extensive and is documented elsewhere (e.g., Carlson <br />and Muth 1989, Minckley and Deacon 1991).. This loss is, for practical purposes, <br />irreversible. The presence of these structures and their role in regulating flows have <br />other, albeit less direct, effects on fish habitat. Structures in the channel may constitute <br /> <br />24 <br /> <br />