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By the time the fish have reached subadult size (250 - 400 mm), they become <br />increasingly piscivorous. Gradually, the subadults begin moving upstream, perhaps <br />drawn to better habitat and more suitable food. Over a period of years, these fish move <br />into adult habitat many miles upstream of the nursery area. Because the movement is <br />so gradual, they become widely distributed in the system. Male Colorado pikeminnow <br />mature when they are about 8 years of age, and females mature when they are about <br />10 years old. <br />~ Razorback Sucker <br />Most of the following text is based on data obtained from studies of razorback suckers <br />in riverine habitat of the UCRB. Despite the fact that most of the razorback suckers <br />remaining in nature are found in Lake Mohave, AZ-NV, rivers are likely to be the focus <br />of restoration efforts in the UCRB. Consequently, the body of information on life <br />~ history, behavior, and habitat use in lacustrine habitat will be used only sparingly. <br />In winter, razorback suckers in the Green River occupy slow runs, slackwaters, eddies, <br />and backwaters of the main channel (McAda and Wydoski 1980, Valdez and Masslich <br />1989). The fish are cold-adapted and remain active in winter; local movements <br />~ increase with increased discharge and flow fluctuations (Valdez and Masslich 1989). <br />Razorback suckers also have been observed using large backwaters in some locations <br />during early spring (Vllestwater Engineering 1996). <br />Razorback suckers in unrestricted riverine habitat exhibit both local and long-distance <br />~ movements in spring and summer (Tyus 1987; Tyus and Karp 1990; Modde et al. <br />1995), although these movements are not as extensive as those observed for the <br />Colorado pikeminnow. During spring migrations, fish may move 50 to 190 km to <br />spawning sites in the Green and Yampa rivers (Tyus and Karp 1990). Radiotracking <br />studies and recaptures of tagged fish on spawning grounds have documented homing <br />~ to specific spawning sites, to which the fish show fidelity. Similar homing movements <br />are known for other catostomids, where at least 95% of the spawning fish migrate to <br />their home stream (see Dence 1948, Werner 1979). Recent studies have implicated <br />olfactory imprinting as the mechanism by which razorback suckers locate spawning <br />areas. Scholz et al. (1992, 1993) confirmed that a burst of thyroxine occurred during <br />the sac-fry stage, immediately before the larvae swam up from the cobbles and entered <br />~ downstream drift. Thyroxine activity in other fishes has been associated with a period <br />of sensitivity in which olfactory imprinting can occur (Hasler and Scholz 1983, Scholz et <br />al. 1992, 1993). <br />The RIP has supported two studies of olfactory imprinting in razorback suckers. Fish <br />~ exposed to two different chemicals during the sac-fry stage were later tested in a maze <br />when they were adults. In the first study, most fish moved and oriented to the correct <br />chemical (Haines et al. 1996). Results were less clear in a subsequent study where <br />fewer fish oriented to either chemical (A. Scholz, personal communication), but <br />behavior may have been confounded by the presence of ripe females. In both studies, <br />19 <br /> <br />