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The exotic zebra mussel, Dreifrena <br />polymorpha, is an unwelcome invader of <br />North American freshwater systems. <br />This species was discovered in Lake St. <br />Clair in 1988, where it presumably was <br />established from discharges of ship <br />ballast water in 1986 (Miller et al. <br />1992; Leach 1993). The freshwater <br />ballast, obtained in an European port, <br />probably discharged veligers which <br />thrived in food-rich Lakes Erie and St. <br />Clair. The zebra mussel spread <br />throughout the Laurentian Great Lakes <br />in the United States and Canada, and <br />into 11 States in the next 4 years <br />(Figure 2). It has been detected in the <br />Mississippi River above and below the <br />mouth of the Missouri River, in the <br />Arkansas River, and has probably found <br />its way into the Missouri River <br />(although undetected as yet). Zebra <br />mussels are expected to spread in the <br />Mississippi River, where their dispersal <br />has been similar to that documented <br />for the Great Lakes (Tucker et al. <br />1993). <br />F/GUi7E Z. ZEBRA MUSSEL S/GHTINGS. <br />Native to the Caspian region of <br />western Russia, the zebra mussel <br />historically inhabited the Aral. <br />Caspian, Azov, and Black seas. The <br />species also was a common inhabitant <br />of the mouths of large rivers that flow <br />into these saline basins (Karnaukhov <br />and Karnaukhov 1993). The zebra <br />mussel began increasing its range in <br />the 19th century by invading North <br />European rivers that emptied into the <br />Black Sea, presumably in response to <br />canal building and increased eutrophi- <br />cation (Karanaukhov and Karanaukhov <br />1993). Europeans have experienced its <br />continuing invasion through shipping <br />canals for nearly 200 years (and more <br />recently in association with hydroelec- <br />tric projects). By about 1930, the <br />mussel had spread throughout Europe <br />and had reached Great Britain. <br />Scientists have speculated (since about <br />1921) that zebra mussels might be <br />transmitted to the United States by <br />shipping (Nalepa and Schlosser 1993), <br />but little was done to reduce the <br />likelihood of such an invasion. <br />