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Once established, the zebra mussel <br />is a prolific colonizer. Maturing within <br />12 to 18 months in favorable environ- <br />ments, it may spawn more than once a <br />year, and production of over 1 million <br />eggs per season is possible from one <br />female (Miller et al. 1992). Fertilized <br />eggs develop into free-swimming <br />larvae (veligers) that can remain <br />suspended in the water for a month. <br />Larvae attach to hard surfaces and <br />transform into the hard-shelled form <br />(post-veligers). Up to 700,000 post <br />veligers/m' have been reported to settle <br />in raw water systems during a single <br />spawning season (Miller et al. 1992; <br />McMahon et al. 1993). In Lake St. <br />Clair, the striped, yellowish-brown <br />mussels grow nearly an inch in the first <br />year, reach a size of 1.5 to 2 inches, and <br />live about 3 years (Mackie 1993). <br />Young zebra mussels settle on various <br />surfaces and firmly attach to them with <br />threads secreted from a gland at the base <br />of the muscular foot (Miller et al. 1992). <br />F/GORE .~. P/PE BLOCKED W/TH ZEBRA MUSSELS. <br />They also can attach to each other, and <br />thus form mussel clusters with <br />population densities of 5,000 to 30,000 <br />mussels/mz not being uncommon. <br />High densities of 342,000 to 700,000 <br />mussels/mZ have been reported (Miller <br />et al. 1992; Leach 1993), and at high <br />densities zebra mussels can form dense <br />mats of 12 inches thick or perhaps <br />more. Because concrete structures, <br />steel pipes, and other man-made items <br />are good attachment substrate for the <br />After its introduction into Lake St. <br />Clair, economic damages caused by the <br />mussel in the Great Lakes was <br />estimated at $5 billion (Nonindigenous <br />Aquatic Nuisance Prevention and <br />Control Act of 1990). Infestations of <br />zebra mussels ranging from 1,000/mZ <br />to 750,000/mZ have been recorded <br />from 10 power plants located from <br />Lake Huron to Lake Erie (Kovalak et <br />al. 1993). Costs to the power industry <br />alone has been estimated at $3.1 billion <br />mussels, they have become a major <br />problem for industrial and municipal <br />water supply facilities. Severe problems <br />to power plants have been caused by <br />zebra mussel colonization inside pipes <br />(Figure 3). <br />over a 10 year period, which includes <br />costs for plant shutdown, new design, <br />and annual maintenance (U.S. Con- <br />gress, Office of Technology Assessment <br />1~~3). Some power plants have <br />expended costs of $250,000 per year <br />