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growth efficiency may have been responsible for observed differences between <br />Colorado squawfish and fathead minnow. <br />The hypothesis that growth efficiency was responsible for the <br />competitive advantage of fathead minnow over Colorado squawfish is consistent <br />with the life-history strategies of the fishes. Colorado squawfish are long <br />lived, have large body size, and delayed reproduction. Fathead minnow are <br />short lived, have small body size, and early reproduction. Implicit in these <br />characteristics is one other correlate of life-history theory: short-.lived <br />species have rapid development compared to long-lived species (Pianka 1970). <br />Faster development implies greater growth efficiency, provided that both <br />species eat similar prey and are exposed to identical environmental <br />conditions. Thus, natural selection may have provided fathead minnow with a <br />competitive advantage over larval stages of Colorado squawfish. <br />Because fathead minnow are not native to the Colorado River Basin, there <br />has been no coevolution with Colorado squawfish to facilitate development of <br />partitioning mechanisms that would allow coexistence. Assuming Colorado <br />squawfish do not become extirpated in the wild, two important implications of <br />the lack of coevolution are related to (1) effects of non-native fishes on <br />habitat selection by young Colorado squawfish and (2) ultimate evolutionary <br />effects of the introduction of non-native fishes into the Colorado River <br />Basin. Mechanisms of nursery-habitat selection used by young Colorado <br />squawfish are unknown, but it is plausible that food availability plays a <br />role. The species that is most efficient at capturing and converting food <br />resources to biomass will be able to persist in an environment for a longer <br />time as resources become scarce (Tilman 1982). For larval Colorado squawfish, <br />the alternative to living in a backwater habitat that has become unsuitable is <br />21 <br />