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<br />of fish in the 125-150 mm TL size classes into the 2005 population. They further postulated that <br />environmental effects were playing a role, and thought that fish of that size were age-2 or the <br />2003 year class. We agree that the likely cause was environmental effects but believe the year- <br />class involved was 2004 not 2003. We believe this because smallmouth bass in this area grow to <br />an average size of about 75 mm TL by the end of their first growing season; that size may even <br />be conservative because upstream populations in the Yampa River where water temperatures are <br />cooler and bass hatch later are of similar size. By the time Christopherson et al. (2005) <br />conducted removal sampling in August 2005, the now age-1 bass hatched in 2004 could easily <br />have grown to 125 to 150 mm TL by August 2005 in the almost two full growing seasons; <br />growth rate estimates from otoliths of smallmouth bass collected in the Yampa River just <br />upstream corroborate growth rates postulated here (pers. comm. P. Martinez, Colorado Division <br />of Wildlife). In 2004, smallmouth bass in that age-1 size class were about 41% of all fish in the <br />population susceptible to electrofishing (based on size frequencies in Fig. 1, Christopherson et <br />al. 2005). Near-complete absence of that generally abundant size-class from the 2005 Whirlpool <br />Canyon population could be the difference between abundances expected in 2005, based on 2004 <br />estimates and removal levels. This, of course, assumes that the abundance and removal <br />estimates of Christopherson et al. 2005 are unbiased. We believe those abundance estimates are <br />essentially correct (no differential bias among years) and the decline shown in 2005 is real, <br />because our own 2005 sampling showed a similar decline in relative abundance for smallmouth <br />bass in Whirlpool Canyon of approximately 50%; we also showed the expected and similar <br />reduction in Lodore Canyon. Further, the lower Yampa River smallmouth bass removal <br />sampling effort in 2005 showed that a large year-class of the 125-150 mm TL fish were present: <br />no large floods or turbidity events were documented there in 2004 based on observations during <br />drift net sampling. Thus, high displacement of age-0 smallmouth bass by a flood or floods in the <br />upper Green River in 2004 may be responsible for a significant reduction in larger bass <br />abundance in 2005 and perhaps a lasting effect into the future. This is important because even <br />relatively substantial removal efforts in most portions of the Upper Colorado River Basin <br />(Hawkins 2005, middle Yampa River; Fuller, 2005 in the lower Yampa) rarely show any <br />29 <br />