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Bestgen et al. 2006). Upstream reaches of Lodore Canyon now have a mix of sizes, including <br />small fish, as reproduction has been noted throughout the canyon and upstream. In Whirlpool <br />Canyon, modal size was lower than upstream and fish in the 151 to 200-mm TL size class or <br />smaller were most abundant (Christopherson et al. 2005; Bestgen et al. 2006). <br />Large numbers of fish, including smallmouth bass, in drift net samples in the Green River <br />upstream of the Yampa River in 2003 and 2004 associated with turbidity events suggested <br />susceptibility to displacement. Such displacement likely also occurred in the Green River <br />downstream of the Yampa River because the Yampa contributed only about 25% of the flow <br />downstream of the confluence during those events. The literature also suggested negative effects <br />of flow increases on early life stages of smallmouth bass (Lukas and Orth 1995, Peterson and <br />Kwak 1999, references therein); Winemiller and Taylor (1982) documented negative effects of <br />turbidity and high flows on disruption of smallmouth bass spawning, nest destruction, and <br />displacement of young. Increased flows levels via dam releases or turbidity events may be a <br />potential controlling mechanisms for early life stages of smallmouth bass in this portion of the <br />river. <br />We believe that displacement or direct mortality of early life stages of smallmouth bass <br />by flow or turbidity events, as noted in 2004 drift net samples, reduced survival of those age-0 <br />fish and subsequently reduced abundance of the same year class of age-1 smallmouth bass in <br />Lodore and Whirlpool canyons in 2005 (Fig. 16). Inexplicable abundance reductions in 2005 <br />were also noted by Christopherson et al. (2005, M. Fuller conducted sampling in The Whirlpool <br />reach), who conducted abundance estimates for smallmouth bass associated with removal efforts <br />in 2004 and 2005 in the Green River in Whirlpool Canyon. They could not explain the reduction <br />in smallmouth bass abundance in 2004 from 8,000 compared to about 3,300 fish in 2005, a <br />decline of 59%, because they only had 31 % removal rate in 2004. Importantly, that decline did <br />not account for recruitment of abundant smaller bass into the 2005 population, which is usually <br />ample enough to replace the bass removed in the subsequent year: age-1 smallmouth bass were <br />about 41 % of the population sampled in 2004. Christoperson et al. (2005) correctly surmised <br />that a large portion of the difference in abundance in 2004 and 2005 was due to low recruitment <br />28 <br />