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. ~ ~~ <br />~'~ ~~ ~ ~ <br />low flows, the fish used slow an fast runs almost exclusively. The change in <br />habitat use without a correspond ng change in relative habitat availability <br />indicates that other factors al o influence habitat selection. These factors <br />could include changes in quasi of physical habitat features such as <br />diversity, depth, dissolved ox gen, etc., or changes in biotic interactions. <br />Osmundson et al. (1995) inter eted the squawfish behavioral changes as <br />reflective of suboptimal cond tions; the behavioral changes demonstrate the <br />ability of the species to mo 'fy their habitat use patterns to temporarily <br />cope with adverse conditions and do not demonstrate habitat preferences under <br />optimum conditions. Osmunds n et al. (1995), therefore, based their flow <br />recommendations on the habit t-selection behavior of adult squawfish observed <br />during the higher somewhat ors natur~l_flow conditions. Under adjusted <br />baseline conditions for the 1975-1993 period of record, summer flows have <br />fallen below recommended levels in 53 months out of 57 months. <br />Winter (November-March): Osmundson et al. (1995) reported that flows during <br />the winter are usually moderate because no water is diverted for irrigation <br />and because additional water is released through upstream dams to increase <br />reservoir storage capacity in anticipation of spring runoff. The relative <br />availability of slow runs and riffles during the winter was very similar to <br />their availability during summer. As in the summer, backwaters, eddies, and <br />pools were the preferred types of habitat in the winter. However, whereas <br />eddies were most preferred in summer, pools were most preferred in winter. <br />Adult squawfish used fewer habitat types overall during winter than during <br />summer. Although fast runs and riffles were used during the summer, they were <br />not used during the winter. The colder water temperatures in winter which <br />cause lower metabolic rates may account for the avoidance of high velocity <br />sites. Absolute area of pools increases as flows decrease and slow runs lose <br />velocity. Because Osmundson et al. (1995) did not sample low flows in the <br />winter, they could not determine <br />o° a~'.wint s. herefore, they recommended winter flows at the <br />pr~,,~`'moderate flow level which maximized weighted area of preferred habitats. <br />~"`~,J~' Under adjusted baseline conditions for the 1975-1993 period of record, winter <br />. ,~~ ' flows have fallen below recommended levels in 34 months out of 95 months. <br />~~ <br />~ ~ Spring (April-July): Osmundson and Kaeding (1989) reported that squawfish use <br />s of low velocity habitats such as backwaters and flooded gravel pits is <br />greatest during the spring runoff. It is believed that squawfish use these <br />habitats during the runoff to escape the high velocity, low temperature flows <br />of the main channel. Because backwaters, flooded gravel pits, and other low <br />velocity habitats are considerably warmer than the main channel during the <br />runoff, these habitats allow squawfish to extend their growing season <br />substantially. The earlier warming of these habitats may also be important in <br />enabling squawfish to reach spawning condition by the time flow and <br />temperature in the main channel are optimum for spawning. Osmundson et al. <br />(1995) reported that the numbers of backwaters and flooded gravel pits <br />increases with increasing spring flows. (Although the number of backwaters <br />eventually decreases as increasing flows co backwaters to sidechannels, <br />the number of other low velocity habitat ikely i creases as increasing flows <br />inundate additional bottomlands.) The a in the magnitude, duration, <br />and frequency of high spring flows, then decreases the quantity-and the <br />duration and frequency of availability f important low velocity, higher <br /> <br />