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21 <br />Some important habitats, such as inundated floodplain depressions used by <br />razorback suckers for spawning, are located outside the channel. Floodplain <br />depressions are principally derived from abandoned main channels, <br />side-channels, backwaters, and meander cutoffs. <br />The creation of complex channel habitat and the formation and eventual <br />abandonment of channel features from which floodplain depressions are formed <br />occur primarily during spring runoff when flows are of sufficient size and <br />duration to cause major changes in channel morphology through significant <br />erosion and deposition of bed and bank materials. The reduction in the <br />magnitude, duration, and frequency of high spring flows has slowed the rate at <br />which channel morphology changes. Consequently, the creation of complex <br />channel habitat and floodplain depressions has slowed. The placement of <br />riprap and other bank stabilization measures and the construction of dikes and <br />levees impede changes in channel morphology and contribute to the slowed <br />creation of complex channel habitat. In addition, the construction of dikes <br />and levees reduces existing channel habitat complexity by causing <br />channelization of the river. Dikes and levees also isolate existing <br />floodplain depressions from the channel during high flows. The slowed <br />creation of complex channel habitats and new floodplain depressions, the <br />reduction of existing channel habitat complexity, and the isolation of <br />existing floodplain depressions have acted to reduce the quantity and quality <br />of important habitat for endangered fishes. <br />Habitat Maintenance <br />Backwaters, used by various life stages of endangered fish, are damaged by the <br />deposition of fine sediments which reduces their depth and consequently their <br />duration and frequency of inundation. Gravel and cobble substrates, used by <br />squawfish for spawning, are damaged by the infiltration of fine sediments. <br />The establishment of vegetation on backwater sediments and on bars further <br />reduces the value of these habitats for endangered fishes. Furthermore, <br />higher flows are required to flush sediments from vegetated backwaters than <br />from unvegetated ones. Osmundson and Kaeding (1991) reported observations <br />that, on the 15-mile reach during the drought years of 1988 to 1990, <br />backwaters were filling in with silt and spring flows were not sufficient to <br />flush out the fine sediment. Also,- they reported that tamarisk colonized sand <br />and cobble bars. The lower frequency of high water years, therefore, <br />decreases the frequency at which silt and sand is flushed from backwaters, <br />fine sediments are flushed from gravel/cobble substrates, and vegetation is <br />scoured from backwaters and bars. As a result, the frequency at which these <br />habitats are suitable for use by endangered fishes has decreased. <br />~~ ~ Seasonal Habitat Availability <br />~ Summer (August-October): Osmundson et al. (1995) reported that availability <br />~ ~ of habitats did not differ significantly between periods of moderate flows and <br />• low flows. Though absolute area of habitat decreases with declining flows, <br />relative area or percent composition of habitat types changes little. <br />However, squawfish habitat use patterns did change. The fish used a greater <br />variety of habitats during moderate flows than during low flows. During <br />moderate flows, the fish used primarily back~:~tcrs, eddies, and pools. During <br />