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1946 <br />JOHNSON ET AL. <br /> 1,200 <br /> - Yartq~a River ~~~`~ <br /> t,000 --- carlander(t988.1877j ~~ <br />~ ~ <br />444 f <br />~ 800 ~ <br /> normum <br />,~ gifts <br />~-.. <br /> ~ <br /> ~ ~ <br /> ~ <br />400 ~ <br />F~ <br />~f - <br />' <br /> j ~ <br />~~ <br />Jf <br />~~' <br />~ <br /> 1 _ <br />`. <br />1 <br />`' ~ ~ <br />~ <br /> 200 ~ `~, calh <br />h <br />mss <br /> ~ smalEmouth <br /> bass <br />Age <br />9 11 13 1b <br />Ficuet=. 2.-Observed growth of channel catfish (1979- <br />1988; Tyus and Nikirk 1990), northern pike (1987-1991; <br />Martinez 1995; Nesler 1995), and smallmouth bass (2003; <br />Martinez 2004) in the Yampa River, Colorado. "Typical" <br />growth trajectories across the species' ranges (Cazlander 1969, <br />1977) are shown for comparison. <br />Results <br />For channel catfish marked for the population <br />estimate (n = 364 fish), the arithmetic mean TL was <br />472 mm (SE = 4.7) and the geometric mean weight was <br />1,101 g; this body size corresponded to an age of 14 <br />years. For northern pike (n = 295 fish), mean TL was <br />560 mm (SE = 9.0) and geometric mean weight was 990 <br />g, corresponding to an age of 4 years. For smallmouth <br />bass (n =1,400 fish), mean TL was 241 mm (SE = 2.2) <br />and geometric mean weight was 290 g, corresponding <br />to an age of 4 years. smallmouth bass were substantially <br />more numerous (N = 267 fish/km; 95% CI = 234-304 <br />fish/km) than northern pike (5 fish/km; 95% CI = 4.3- <br />6.1 fish/km) or channel catfish (68 fish/km; 95% CI = <br />34-166 fish/km). Channel catfish from the Yampa <br />River grew considerably more slowly than the national <br />average (Carlander 1969), and this disparity increased <br />with age (Figure 2). By age 8, channel catfish were <br />approximately 200 mm smaller than the length at age <br />reported for channel catfish elsewhere. The growth of <br />northern pike as reported by Nesler (1995) was similar <br />to the national average (Carlander 1969), as was the <br />growth of smallmouth bass (Carlander 1977). In 2003 <br />and 2004, mean W was 118 for channel catfish, 93 for <br />northern pike, and 102 for smallmouth bass. <br />The channel catfish diet (n = 32 stomachs examined; <br />30 were. nonempty) included very few fish (0.2% by <br />mass); only 6.6% of stomachs with food contained any <br />fish remains. Most of the diet consisted of virile <br />crayfish Orconectes virilis (53.7%) and plant matter <br />(36.0%); insects (primarily Ephemeroptera, Trichop- <br />tera, and Plecoptera) made up the remainder of the diet <br />(10.1%). Northem pike (n = 45 stomachs; 33 were <br />nonempty) were primarily piscivorous; fish constituted <br />72.2% of the diet (by mass). Northem pike also ate <br />crayfish (24.5%) and a few insects (3.3%). Northem <br />pike preyed nearly equally on catostomids, centrarch- <br />ids, cyprinids, and salmonids. smallmouth bass (n = <br />178 stomachs; 149 were nonempty) consumed mainly <br />crayfish (51.5% by mass). Aquatic insects, including <br />Ephemeroptera (80% by number), Plecoptera (15%), <br />Hemiptera (Corixidae and Notonectidae: 3%), and <br />Trichoptera (<1%), made up 42.8% of the diet. Only <br />5.7% of the diet consisted of fish; because of the <br />advanced state of digestion in the samples, most fish <br />remains could not be definitively identified below the <br />family level. Small cyprinids and smallmouth bass each <br />constituted 40% of the fish prey, and suckers and <br />sunfish made up the remainder. In contrast, fish <br />(minnows and suckers) were the primary prey <br />(67.7%) of smallmouth bass collected from the upper <br />Colorado River (n = 325 stomachs), while crayfish <br />(12.9%) and insects (19.4%) made up much smaller <br />proportions of the diet than was observed in the Yampa <br />River. Northern pike and smallmouth bass consumed <br />fish that were similar in size relative to predator size. <br />The median prey :predator size ratio (•I'L) of fish in <br />stomachs was 0.22 for northern pike and 0.21 for <br />smallmouth bass. Northem pike consumed proportion- <br />ately larger prey (p95 = 0.50) than smallmouth bass (p95 <br />= 0.39), but the minimum prey size consumed was <br />similar between the two species (PS = 0.12 for northern <br />pike and 0.11 for smallmouth bass). <br />Per-capita consumption of fish was highest for <br />northern pike (2.7 kg/year; Table 2). Fish constituted <br />a small fraction of the diet in channel catfish <br />simulations; thus, consumption of fish by the average <br />channel catfish was predicted to be only 3.3 g/year. <br />Per-capita consumption of fish by smallmouth bass was <br />estimated at 0.06 kg/year. At the population level, <br />realized fish consumption by smallmouth bass .(mean = <br />15.2 kg •km-' •year'; 95% CI = 13.3-17.1 <br />kg • km-' •year ') was similar to fish consumption by <br />northern pike (mean = 13.7 kg• km ' •year'; 95% CI <br />= 11.4-16.0 kg • km-' •year ') and was about 65 times <br />higher than fish consumption by channel catfish (mean <br />= 0.22 kg • km ' •year' ; 95% CI = 0.05-0.40 <br />kg• km-' •year '). <br />Potential piscivory by channel catfish was not <br />different from realized consumption, because we <br />assumed that no diet shift would occur. Potential <br />piscivory by the smallmouth bass population was 168.5 <br />kg•km-' •year-' (95% CI = 147.0-189.9 <br />kg • km-' •year '), or about 10 times higher than that <br />by the northern pike population (17.2 kg • km-' •year '; <br />95% CI = 14.2-20.1 kg• km-' •year'). Combined <br />