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Spawning ar,d Egg Survival <br />Little is known about natural reproduction of humpback chubs <br />partly because the species spawn during spring runoff when sampling <br />conditions are poor {Valdez et al. 1~J82). Humpback chub spawning <br />sites have not been located (L. Kaeding, USFWS, pers. comm.}. The <br />species-may spawn in pockets of rubble and gravel within Westwater <br />Canyon and Black Rocks (Miller et al. 1982c) or on submerged cobble <br />bars near the upstream end of Brlack Rocks (Valdes and Nilson 1982). <br />Some humpback chubs may spawn just outside the deepwater areas of <br />Black Rocks (Archer et al. 1985). <br />Humgbaek chubs in the Little Colorado River probably spawned <br />during April and May in 1981 (Kaeding and Zimmerman 1983).. <br />Suspected spawning at Black-Rocks in 1981 to 1984 ranged from <br />. mid-May to end of July, and water temperatures ranged from 11.5 C to <br />23 C (Valdez and Nilson 1982, Archer et al.. 1985). Optimum egg <br />survival is at 20 C (Bullcley et al. 1982, Marsh 1985). Incidence of <br />abnormal fry is lowest at 2D C (Marsh 1985), and survival of fry is <br />greatest at 21-22 C (Hamman 1982). <br /> <br />Reasons for Decline and Limiting Factors <br />Decline of humpback chub populations is attributed mainly to <br />loss of Habitat caused by construction and operation of Flaming <br />Gorgcr and (;1Fn Canyon dams, and perh~~ps Hoover Dam (liSFlti'S 1J7J). <br />Reservoic•s and their cold tailwaters are nut suitable for <br />9 <br />