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20001 XYRAUCHEN TEXANUS IN LAKE MOHAVE 183 <br />TABLE 1. Individual tracking summaries for sonic telemetry of 10 wild adult Xyrauchen texanus in Lake Mohave, Ari- <br />zona and Nevada, November 1994 to July 1997. Fish are individually identified by tag code; number of observations, <br />average and total distance (m d-t) traveled, days tracked, and range (km) of each fish are presented. <br /> <br /> <br />Fish <br />ID <br /> <br />TL <br />(mm) <br /> <br />Weight <br />(kg) <br /> <br />No. of <br />obs. <br />Distance traveled <br />Min. Max. Avg. <br />- - - - - - (m d-t) - - - - - - - <br /> <br />s Total <br />distance <br />traveled <br />(km) <br /> <br />Days <br />tracked <br /> <br />Range <br />(WPa) <br />FEMALES <br />88 615 2.70 65 0 6,703 603 1322 250.4 974 32-66 <br />276 654 3.61 71 21 3,502 606 708 352.5 953 24-67 <br />339 665 3.33 62 9 10,108 767 1413 417.4 981 27-99 <br />375 680 3.35 54 4 2,368 417 544 249.7 540 16-64 <br />384 631 2.76 36 0 6,886 614 1133 18310 466 26-65 <br />465 614 2.94 65 9 6,558 485 851 2643 969 5-34 <br />x 643 3.12 59 7 6,021 582 996 286.2 814 <br />MALES <br />2228 558 2.40 51 3 15,607 788 2280 231.5 540 39-90 <br />366 588 2.29 57 0 1,986 243 509 167.7 974 31-62 <br />348 526 1.75 65 0 17,353 509 2154 181.7 960 32-71 <br />258 595 2.26 57 0 1,635 242 334 120.9 540 29-47 <br />X 567 118 58 1 9,145 446 1319 175.5 754 <br />eWP = way-point <br />where cold Hoover Dam releases mix with <br />warmer Lake Mohave waters. We observed 9 <br />X. texanus (nontagged) near the surface at the <br />mixing zone on 8 June 1997 (9 fish per 1.5 <br />km). <br />Movements <br />Fish movements were variable and distinc- <br />tive by sex and among individuals. Both males <br />(> 17.4 km d-1) and females (10.1 km d-1) trav- <br />eled substantial distances in short periods of <br />time, while some fish remained sedentary for <br />days. Fish on average moved 527 in d-1 (u = <br />242-788 in d-I), and while there was no statis- <br />tical difference between sexes, females tended <br />to be slightly more active (ac = 582 in d-I) <br />than males (x = 446 in d-1; Table 1). Hourly <br />movement rates were statistically examined, <br />and although no significant trends were iden- <br />tified, apparently fish might be slightly more <br />active during summer at midmorning hours <br />(0900-1000 h), supporting previous observa- <br />tions (McAda and Wydoski 1980). <br />Movement rates were examined both for <br />nondirectional (m d-I) and directional (linear <br />km month-1) activities. Monthly averages near <br />the end of, or shortly after, spawning (March, <br />April, May) exceeded 1000 in d-1 for 1995, <br />1996, and 1997 (Table 2). Linear distances <br />traveled (km month-1) peaked following <br />spawning for all 3 yr (March 1995, May 1996, <br />April 1997 [Table 2]). Linear ranges for indi- <br />viduals (N = 10) averaged 39 km (18-72 km) <br />during the course of the study. Females on <br />average ranged 42 km (29-72 km) compared <br />to 35 km (18-51 km) for males. <br />All study fish returned to spawning sites <br />used in previous years, but they also visited <br />other spawning sites. Spawning (31 Janu- <br />ary-15 March 1995, 1996) females were signif- <br />icantly (ANOVA/Fl g = 6.79, P = 0.0314) more <br />active (460 vs. 87 m d-1) than males. During <br />peak spawning (1-28 February 1995, 1996), <br />the majority of females crossed the reservoir <br />to visit other spawning sites while males typi- <br />cally remained at a single site (Fig. 2). <br />Habitat Use <br />Xyrauchen texanus exhibited a significant <br />association with inshore (<50 in of shore) habi- <br />tats (P < 0.001, chi square test). This affinity <br />was most pronounced during spawning when <br />distance from shore averaged <30 in. Fish <br />moved offshore during summer (July-Novem- <br />ber), possibly to avoid warmer water tempera- <br />tures (Fig. 3). <br />During warmer months the reservoir pre- <br />sented fish a wide range of thermally stratified <br />habitats. Temperatures up-reservoir were cold <br />(12°-16°C), regardless of season or depth, due <br />to hypolimnetic releases from Lake Mead. <br />During summer the reservoir thermally strati- <br />fies further downstream presenting tempera- <br />tures ranging from 30°C (surface) to 15°C