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182 WESTERN NORTH AMERICAN NATURALIST [Volume 60 <br />Survey Design <br />We conducted weekly surveys by boat <br />using a global positioning system (GPS) and a <br />preestablished survey grid composed of 71 <br />way-points or "listening stations" programed <br />into a CPS navigational system (Fig. 1). Sites <br />were at <1600-m intervals to standardize <br />search effort and ensure systematic coverage <br />of the reservoir. Transmitter signals were <br />detected using a DH-2 hand-held, directional <br />hydrophone, a USR-5W ultrasonic receiver, <br />and headphones. When signals were detected <br />from way-points, we searched out the exact <br />location and further triangulated using the <br />hydrophone and GPS. A digital readout pro- <br />vided time interval measurements between <br />signals that correlated to tag (fish) tempera- <br />ture. Recorded data included fish location (lat- <br />itude and longitude), body temperature, and <br />estimated distance (<10 in, 10-50 in, 50-100 m, <br />> 100 m) from shore. Vertical water tempera- <br />ture profiles were measured using a Hydro- <br />labTM temperature profiler between May and <br />October 1995. <br />Surveys were scheduled biweekly for the <br />14-month transmitter life and were conducted <br />in 2 shifts: 0200-1000 h and 1400-2200 h. <br />Transmitter life greatly exceeded manufac- <br />turer estimates, allowing us to collect addi- <br />tional intermittent information between April <br />and August 1996 and from April through July <br />1997, <br />was determined through observations and the <br />collection of newly emergent larval X. texanus <br />by an independent program (Mueller 1995). <br />Movement rates (m d-I) during active spawn- <br />ing (31 January-15 March 1995, 1996) for both <br />sexes were examined for differences in move- <br />ment patterns and distances using an analysis <br />of variance (P < 0.05; Sokal and Rohlf 1981). To <br />compare male and female movement patterns <br />when spawning observations and larval sucker <br />densities were most prevalent, we mapped <br />individual fish movement during peak spawn- <br />ing (1-28 February). <br />RESULTS <br />A total of 20 X. texanus (10 males and 10 <br />females) were surgically implanted with sonic <br />transmitters. Six females and 4 males were <br />tracked during the entire 14-month study <br />period and were used in this analysis (Table 1; <br />see Mueller et al. 1998). Remaining fish were <br />excluded from analysis because either we lost <br />contact during the study period or transmit- <br />ters became immobile (shed tag or mortality). <br />An expanded description can be found in <br />Mueller et al. (1998). Study fish averaged 609 <br />mm (555-680 mm) total length and 2.78 kg <br />(1.75-4.21 kg). More than 1000 contacts were <br />recorded representing 580 daily fish sightings, <br />300 hourly sightings, 800 body temperatures, <br />and 100 vertical water temperature profiles. <br />Data Analysis <br />Using digitized shoreline profile maps of <br />Lake Mohave and the geographical informa- <br />tion system (GIS) program ARCINFOTM, we <br />created movement maps and calculated dis- <br />tances (m) and movement rates (m d-I) be- <br />tween detection locations. Rate measurements <br />and distances from shore were transformed <br />into single classification frequency distribu- <br />tions, and observed data were summarized by <br />classes. Shoreline use was examined by com- <br />paring percentiles of available surface habitat <br />to the percentile of fish use within specific <br />zones via a chi square test (P < 0.001; Sokal <br />and Rohlf 1981). Comparisons of shoreline habi- <br />tat utilization, movement rates, and area affin- <br />ity were divided into spawning (November- <br />April) and nonspawning (May-October) peri- <br />ods as defined by reports for X. texanus in <br />Lake Mohave (Minckley 1983, Bozek et al. <br />1990, Minckley et al. 1991). Active spawning <br />Distribution <br />Fish distribution was distinctly seasonal. <br />Some areas of the reservoir were occupied <br />year-round, while other locations were used <br />specifically for spawning or as summer sanctu- <br />aries. Big Basin and Owl Point areas (Fig. 1) <br />were used year-round; Tequila and Yuma coves <br />and the warm/cold mixing zone were used <br />seasonally. Fish were seldom detected in up- <br />or down-reservoir canyon habitats. Detection <br />numbers were highest between way-point <br />(WP) 30 and WP 37. Fish found in Big Basin <br />typically were along the Nevada shoreline be- <br />tween Nine Mile and Six Mile coves (Fig. 2). <br />This area was used by 3 fish during spawning <br />and by 5 during summer. Tequila Cove (WP 39) <br />was visited by all but 1 fish during spawning. <br />Owl Point (WP 32) was visited by all <br />spawners and by 4 fish during summer and <br />autumn. Following spawning, 7 suckers moved <br />into the area between WP 12 and WP 24