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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />3.1.1. Tyus and Karp (1989) <br /> <br />The authors evaluated habitat use and streamflow needs of the Colorado pikeminnow, humpback <br />chub, bonytail and razorback sucker in the Yampa River based on a compilation of existing data, <br />both publish and unpublished. They examined the habitat requirements and factors limiting the <br />distribution and abundance of each species by life stage. Although they discussed year-round <br />flow needs in general terms, they did not quantify these flow needs. Major conclusions drawn <br />from this investigation are: <br /> <br />1. the Yampa and Green rivers constitute the most important riverine-system for the <br />maintenance and recovery of endangered fishes; <br /> <br />2. flows in the Yampa provide a natural shape to the hydrograph of both rivers, mitigating <br />the effects on native fishes due to flow regulation upstream on the Green; <br /> <br />3. high spring flows may inhibit invasion and proliferation of nonnative fishes; <br /> <br />4. Colorado pikeminnow and razorback suckers rely on habitats in both rivers to meet <br />various life stage requirements; <br /> <br />5. therefore, these two river basins should be considered as a single ecosystem when <br />determining the needs of the endangered fishes. <br /> <br />Each of the major components of the annual hydrograph plays a role in the life history of the <br />native fishes. Any significant modification of the hydrograph poses an obstacle to recovering the <br />endangered fishes. However, additional studies are needed to better quantify flow needs. <br /> <br />3.1.2. Harvey, Mussetter and Wick (1993) <br /> <br />Several spawning sites for the Colorado pikeminnow have been identified on cobble-gravel bars <br />in the lower Yampa Canyon. The authors examined the physical processes that create and <br />maintain these spawning bars during the 1991 and 1992 spring runoff seasons. <br /> <br />Pikeminnow begin migrating to these sites in May and early June and spawn on the recessional <br />limb of the spring hydrograph, when water temperatures typically are between 17 and 2rC. <br />Timing of spawning varies with the magnitude of flow, occurring earlier in low-water years and <br />later in high-water years. Pikeminnow require a clean gravel substrate on which to deposit their <br />eggs, as well as relatively quiet resting-staging areas in adjacent pools or shoreline eddies. <br /> <br />High spring peak flows are known to form and transform the major morphological features of the <br />river channel, including mid-channel cobble-gravel bars that provide spawning habitat for the <br />pikeminnow in Yampa Canyon. Sediments are entrained and conveyed by higher stream flows. <br />Velocity, rather than volume, is the critical factor that determines the capacity of the stream to <br />move sediment. Finer sediments (silts and sands) are more easily mobilized, while coarser <br />sediments (gravels and cobbles) require higher velocities to move them. Harvey et al. found that <br />as velocity increases on the ascending limb of the spring hydrograph, sediment is transported <br />from upstream sources to the vicinity of the spawning ITc!rs. The Little Snake River is a <br />significant source of this sediment. However, during the highest flows (19,500-27,500 cfs), <br />heterogeneous, poorly sorted sediments (silts to cobbles) are deposited in backwaters created by <br />sharp bends in the sandstone-embedded channel. These deposits are unsuitable for spawning. <br />As spring flows recede, finer sediments are scoured from the interstices of the coarser gravels <br />and cobbles to provide a suitable substrate for spawning. Thus, the entire springhydrograph is <br />necessary to transport, deposit, shape and clean spawning substrates for pikeminnow. <br /> <br />9 <br />