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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />2 <br /> <br />Similar to most Colorado River Basin rivers, the fish fauna of the Yampa River is dominated by <br />cyprinids and catostomids. A total of 12 fishes are native to the Yampa River, and at least 20 nonnative <br />fish species have been introduced (fable 1). The upper Yampa River and i~ tnbutaries are typical trout <br />waters, and a transition to cool- or warmwater fishes occurs at lower elevations. Of the native fishes, seven <br />are considered in jeopardy or declining and have federal status under the 1973 Endangered Species Act as <br />amended; four are endangered and three are under review (candidates) for federal listing. Most of the <br />nonnative fishes have established reproducing populations. <br />Fish collections used in our analyses were from the lower 94.1 km (river km) of the Yampa River, <br />between the upper end of Cross Mountain Canyon and the Yampa-Green River confluence (Figure 1). <br />This section of river was divided into five reaches defined mainly by prominent changes in terrain and river <br />morphology (Figures 1, 2). Cross Mountain Canyon (Reach 5) is a high-gradient reach with boulder, <br />cobble, and gravel substrates. Mainchannel habitat is primarily rocky falls, rapids, and runs. Small <br />shoreline pools and eddies occur behind boulders. Reach 4 is a low- to moderate-gradient stretch with <br />cobble, gravel, and sand substrates. Terrain is open and hilly. Mainchannel habitat mostly follows a <br />repetitive run-rime-pool sequence, and shoreline habitat consists primarily of pools and embayments. <br />From the lower end of Lily Park downstream to Deerlodge Park (Reach 3), the Yampa River broadens <br />and slowly meanders through flat, open terrain and braids around sand and gravel bars. Numerous <br />backwaters and shoreline embayments occur in this reach. Substrates are primarily gravel, sand, and silt <br />The lower 72.2 km of the Yampa River are contained within Yampa Canyon. In upper Yampa Canyon <br />(Reach 2), the Yampa River is mostly a high-gradient system of rocky runs and rapids, eddies, and riffles. <br />Boulder, cobble, and gravel substrates predominate. Downstream of Harding Hole to the Yampa-Green <br />River confluence Qower Yampa Canyon, Reach I), channel gradient is mostly low to moderate, and <br />mainchannel habitat consists primarily of deep runs and pools and braided sections interspersed with gravel <br />and cobble bars. In reaches 1 and 2, shoreline habitat is mostly pools, eddies, and embayments. Extent of <br />backwater development is higher in Reach 1 than in Reach 2. <br /> <br />MEmODS <br /> <br />SampUng <br /> <br />The main objective of fish sampling conducted in the lower 94.1 km of the Yampa River during <br />1980-1984 was to document Colorado squawflSh reproduction through collection of age-O fish. Accordingly, <br />annual sampling schedules were based primarily on knowledge of timing and location of Colorado <br />squawfish spawning in the Upper Colorado River System, particularly the Green River sub-system, and <br />were revised as new information was produced. Vanicek and Kramer (1969) collected adult Colorado <br />squawfish in reproductive condition (ripe) from the Green River, Utah, in late June through early August, <br />1964-1966. They reported that, in all years, ripe Colorado squawflSh were first collected about 1 month <br />