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<br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br />I <br /> <br />r <br />f' <br />" <br />~. <br />~.. <br />if <br />f <br /> <br />14 <br /> <br />largest numbers occurring in reservoirs, from where they moved or were carried to lotic <br /> <br />habitats. Unlike canyon-bound reaches or narrow, bedrock-constrained channels of <br /> <br />smaller low desert streams exposed to violent flooding which reduced nonnative <br /> <br />populations, flooding in the low-gradient, lower Colorado River may have had little affect <br /> <br />on non-native species (see Minckley and Meffe 1987). Minckley (1991) considered <br /> <br />predatory, competitive, and other biotic pressures of non-native on native fishes the most <br /> <br />important factors now forcing native biotas of western North America toward extinction <br /> <br />(see also, contributions in Minckley and Deacon 1991). <br /> <br /><. <br />~,;-. <br /> <br />3. Study Organism, the Razorback Sucker <br />I studied the razorback sucker, a large-bodied, long-lived, <br /> <br />iteroparous species endemic to the Colorado River system of western North America <br /> <br /> <br />(Minckley 1973, 1983), as a species adapted to conditions in a large, highly seasonal <br /> <br /> <br />river. To successfully recruit in the natural state, I assume this species depends on cues <br /> <br /> <br />and conditions provided by fluctuating seasonal discharge, as with other large fishes in <br />large-river habitat (Lowe-McConnell 1987; Junk et al. 1989; Winemiller 1989; Schlosser <br />1991; Gherke 1992). <br /> <br /> <br />Closely spaced gill-rakers and a sub-terminal mouth suggest adaptations for <br /> <br />feeding in a benthic environment (Hubbs and Miller 1953; Minckley et al. 1991). In <br /> <br /> <br />reservoirs adult diets consist largely of filtered zooplankton which they collect in mid- <br /> <br /> <br />water (Roden 1978; Marsh 1987). In the river, mainstem, eddy or backwater habitats, <br /> <br /> <br />diet consists mainly of benthic material composed of chironomids and detritus. <br />